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Hearing range describes the range of
frequencies Frequency is the number of occurrences of a repeating event per unit of time. It is also occasionally referred to as ''temporal frequency'' for clarity, and is distinct from ''angular frequency''. Frequency is measured in hertz (Hz) which is e ...
that can be heard by humans or other animals, though it can also refer to the range of levels. The human range is commonly given as 20 to 20,000 Hz, although there is considerable variation between individuals, especially at high frequencies, and a gradual loss of sensitivity to higher frequencies with age is considered normal. Sensitivity also varies with frequency, as shown by
equal-loudness contour An equal-loudness contour is a measure of sound pressure level, over the frequency spectrum, for which a listener perceives a constant loudness when presented with pure steady tones. The unit of measurement for loudness levels is the phon and ...
s. Routine investigation for hearing loss usually involves an audiogram which shows threshold levels relative to a normal. Several animal species are able to hear frequencies well beyond the human hearing range. Some dolphins and bats, for example, can hear frequencies over 100 kHz. Elephants can hear sounds at 14–16 Hz, while some whales can hear infrasonic sounds as low as 7 Hz.


Measurement

A basic measure of hearing is afforded by an audiogram, a graph of the
absolute threshold of hearing The absolute threshold of hearing (ATH) is the minimum sound level of a pure tone that an average human ear with normal hearing can hear with no other sound present. The absolute threshold relates to the sound that can just be heard by the organi ...
(minimum discernible sound level) at various frequencies throughout an organism's nominal hearing range. Behavioural hearing tests or physiological tests can be used to find hearing thresholds of humans and other animals. For humans, the test involves tones being presented at specific frequencies ( pitch) and intensities (
loudness In acoustics, loudness is the subjective perception of sound pressure. More formally, it is defined as, "That attribute of auditory sensation in terms of which sounds can be ordered on a scale extending from quiet to loud". The relation of ph ...
). When the subject hears the sound, they indicate this by raising a hand or pressing a button. The lowest intensity they can hear is recorded. The test varies for children; their response to the sound can be indicated by a turn of the head or using a toy. The child learns what to do upon hearing the sound, such as placing a toy man in a boat. A similar technique can be used when testing animals, where food is used as a reward for responding to the sound. The information on different mammals hearing was obtained primarily by behavioural hearing tests. Physiological tests do not need the patient to consciously respond.


Humans

In humans, sound waves funnel into the ear via the external ear canal and reach the
eardrum In the anatomy of humans and various other tetrapods, the eardrum, also called the tympanic membrane or myringa, is a thin, cone-shaped membrane that separates the external ear from the middle ear. Its function is to transmit sound from the ...
(tympanic membrane). The compression and
rarefaction Rarefaction is the reduction of an item's density, the opposite of compression. Like compression, which can travel in waves ( sound waves, for instance), rarefaction waves also exist in nature. A common rarefaction wave is the area of low relat ...
of these waves set this thin membrane in motion, causing sympathetic vibration through the middle ear bones (the
ossicles The ossicles (also called auditory ossicles) are three bones in either middle ear that are among the smallest bones in the human body. They serve to transmit sounds from the air to the fluid-filled labyrinth (cochlea). The absence of the auditor ...
: malleus, incus, and stapes), the basilar fluid in the cochlea, and the hairs within it, called
stereocilia Stereocilia (or stereovilli or villi) are non-motile apical cell modifications. They are distinct from cilia and microvilli, but are closely related to microvilli. They form single "finger-like" projections that may be branched, with normal cell ...
. These hairs line the cochlea from base to apex, and the part stimulated and the intensity of stimulation gives an indication of the nature of the sound. Information gathered from the hair cells is sent via the auditory nerve for processing in the brain. The commonly stated range of human hearing is 20 to 20,000 Hz.20 to 20,000 Hz corresponds to sound waves in air at 20°C with
wavelength In physics, the wavelength is the spatial period of a periodic wave—the distance over which the wave's shape repeats. It is the distance between consecutive corresponding points of the same phase on the wave, such as two adjacent crests, tr ...
s of 17 meters to 1.7 cm (56 ft to 0.7 inch).
Under ideal laboratory conditions, humans can hear sound as low as 12 Hz and as high as 28 kHz, though the threshold increases sharply at 15 kHz in adults, corresponding to the last auditory channel of the
cochlea The cochlea is the part of the inner ear involved in hearing. It is a spiral-shaped cavity in the bony labyrinth, in humans making 2.75 turns around its axis, the modiolus. A core component of the cochlea is the Organ of Corti, the sensory o ...
. The human auditory system is most sensitive to frequencies between 2,000 and 5,000 Hz. Individual hearing range varies according to the general condition of a human's ears and nervous system. The range shrinks during life, usually beginning at around the age of eight with the upper frequency limit being reduced. Women lose their hearing somewhat less often than men. This is due to a lot of social and external factors. For example, men spend more time in noisy places, and this is associated not only with work but also with hobbies and other activities. Women have a sharper hearing loss after menopause. In women, hearing decrease is worse at low and partially medium frequencies, while men are more likely to suffer from hearing loss at high frequencies. Audiograms of human hearing are produced using an audiometer, which presents different frequencies to the subject, usually over calibrated headphones, at specified levels. The levels are
weighted A weight function is a mathematical device used when performing a sum, integral, or average to give some elements more "weight" or influence on the result than other elements in the same set. The result of this application of a weight function is ...
with frequency relative to a standard graph known as the minimum audibility curve, which is intended to represent "normal" hearing. The threshold of hearing is set at around 0 
phon The phon is a logarithmic unit of loudness level for tones and complex sounds. Loudness is measured in sone which is a linear unit. Human sensitivity to sound is variable across different frequencies; therefore, although two different tones may ...
on the equal-loudness contours (i.e. 20 micropascals, approximately the quietest sound a young healthy human can detect),Gelfand, S A., 1990. ''Hearing: An introduction to psychological and physiological acoustics''. 2nd edition. New York and Basel: Marcel Dekker, Inc. but is standardised in an
ANSI The American National Standards Institute (ANSI ) is a private non-profit organization that oversees the development of voluntary consensus standards for products, services, processes, systems, and personnel in the United States. The organi ...
standard to 1 kHz. Standards using different reference levels, give rise to differences in audiograms. The ASA-1951 standard, for example, used a level of 16.5  dB SPL (sound pressure level) at 1 kHz, whereas the later ANSI-1969/ISO-1963 standard uses , with a 10 dB correction applied for older people.


Other primates

Several
primates Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter including ...
, especially small ones, can hear frequencies far into the ultrasonic range. Measured with a signal, the hearing range for the
Senegal bushbaby The Senegal bushbaby (''Galago senegalensis''), also known as the Senegal galago, the lesser galago or the lesser bush baby, is a small, nocturnal primate, a member of the galago family Galagidae. The name "bush baby" may come either from the a ...
is 92 Hz–65 kHz, and 67 Hz–58 kHz for the
ring-tailed lemur The ring-tailed lemur (''Lemur catta'') is a large strepsirrhine primate and the most recognized lemur due to its long, black and white ringed tail. It belongs to Lemuridae, one of five lemur families, and is the only member of the ''Lemur'' ...
. Of 19 primates tested, the Japanese macaque had the widest range, 28 Hz–34.5 kHz, compared with 31 Hz–17.6 kHz for humans.


Cats

Cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
s have excellent hearing and can detect an extremely broad range of frequencies. They can hear higher-pitched sounds than humans or most dogs, detecting frequencies from 55  Hz up to 79  kHz. Cats do not use this ability to hear
ultrasound Ultrasound is sound waves with frequencies higher than the upper audible limit of human hearing. Ultrasound is not different from "normal" (audible) sound in its physical properties, except that humans cannot hear it. This limit varies ...
for communication but it is probably important in hunting, since many species of rodents make ultrasonic calls. Cat hearing is also extremely sensitive and is among the best of any mammal, being most acute in the range of 500 Hz to 32 kHz. This sensitivity is further enhanced by the cat's large movable outer ears (their '' pinnae''), which both amplify sounds and help a cat sense the direction from which a noise is coming.


Dogs

The hearing ability of a dog is dependent on breed and age, though the range of hearing is usually around 67 Hz to 45 kHz. As with humans, some dog breeds' hearing ranges narrow with age, such as the
German shepherd The German Shepherd or Alsatian is a German breed of working dog of medium to large size. The breed was developed by Max von Stephanitz using various traditional German herding dogs from 1899. It was originally bred as a herding dog, for ...
and miniature poodle. When dogs hear a sound, they will move their ears towards it in order to maximize reception. In order to achieve this, the ears of a dog are controlled by at least 18 muscles, which allow the ears to tilt and rotate. The ear's shape also allows the sound to be heard more accurately. Many breeds often have upright and curved ears, which direct and amplify sounds. As dogs hear higher frequency sounds than humans, they have a different acoustic perception of the world. Sounds that seem loud to humans often emit high-frequency tones that can scare away dogs.
Whistle A whistle is an instrument which produces sound from a stream of gas, most commonly air. It may be mouth-operated, or powered by air pressure, steam, or other means. Whistles vary in size from a small slide whistle or nose flute type to a lar ...
s which emit ultrasonic sound, called
dog whistle A dog whistle (also known as silent whistle or Galton's whistle) is a type of whistle that emits sound in the ultrasonic range, which humans cannot hear but some other animals can, including dogs and domestic cats, and is used in their trainin ...
s, are used in dog training, as a dog will respond much better to such levels. In the wild, dogs use their hearing capabilities to hunt and locate food. Domestic breeds are often used to guard property due to their increased hearing ability. So-called "Nelson"
dog whistle A dog whistle (also known as silent whistle or Galton's whistle) is a type of whistle that emits sound in the ultrasonic range, which humans cannot hear but some other animals can, including dogs and domestic cats, and is used in their trainin ...
s generate sounds at frequencies higher than those audible to humans but well within the range of a dog's hearing.


Bats

Bats have evolved very sensitive hearing to cope with their nocturnal activity. Their hearing range varies by species; at the lowest it can be 1 kHz for some species and for other species the highest reaches up to 200 kHz. Bats that can detect 200 kHz cannot hear very well below 10 kHz. In any case, the most sensitive range of bat hearing is narrower: about 15 kHz to 90 kHz. Bats navigate around objects and locate their prey using echolocation. A bat will produce a very loud, short sound and assess the echo when it bounces back. Bats hunt flying insects; these insects return a faint echo of the bat's call. The type of insect, how big it is and distance can be determined by the quality of the echo and time it takes for the echo to rebound. There are two types of call constant frequency (CF), and
frequency modulated Frequency modulation (FM) is the encoding of information in a carrier wave by varying the instantaneous frequency of the wave. The technology is used in telecommunications, radio broadcasting, signal processing, and computing. In analog freq ...
(FM) that descend in pitch. Each type reveals different information; CF is used to detect an object, and FM is used to assess its distance. The pulses of sound produced by the bat last only a few thousandths of a second; silences between the calls give time to listen for the information coming back in the form of an echo. Evidence suggests that bats use the change in pitch of sound produced via the
Doppler effect The Doppler effect or Doppler shift (or simply Doppler, when in context) is the change in frequency of a wave in relation to an observer who is moving relative to the wave source. It is named after the Austrian physicist Christian Doppler, who ...
to assess their flight speed in relation to objects around them. The information regarding size, shape and texture is built up to form a picture of their surroundings and the location of their prey. Using these factors a bat can successfully track change in movements and therefore hunt down their prey.


Mice

Mice A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus'' ...
have large ears in comparison to their bodies. They hear higher frequencies than humans; their frequency range is 10 kHz to 70 kHz. They do not hear the lower frequencies that humans can; they communicate using high-frequency noises some of which are inaudible by humans. The distress call of a young mouse can be produced at 40 kHz. The mice use their ability to produce sounds out of predators' frequency ranges to alert other mice of danger without exposing themselves, though notably, cats' hearing range encompasses the mouse's entire vocal range. The squeaks that humans can hear are lower in frequency and are used by the mouse to make longer distance calls, as low-frequency sounds can travel farther than high-frequency sounds.


Birds

Hearing is birds' second most important sense and their ears are funnel-shaped to focus sound. The ears are located slightly behind and below the eyes, and they are covered with soft feathers – the auriculars – for protection. The shape of a bird's head can also affect its hearing, such as owls, whose facial discs help direct sound toward their ears. The hearing range of birds is most sensitive between 1 kHz and 4 kHz, but their full range is roughly similar to human hearing, with higher or lower limits depending on the bird species. No kind of bird has been observed to react to ultrasonic sounds, but certain kinds of birds can hear infrasonic sounds. "Birds are especially sensitive to pitch, tone and rhythm changes and use those variations to recognize other individual birds, even in a noisy flock. Birds also use different sounds, songs and calls in different situations, and recognizing the different noises is essential to determine if a call is warning of a predator, advertising a territorial claim or offering to share food." "Some birds, most notably oilbirds, also use echolocation, just as bats do. These birds live in caves and use their rapid chirps and clicks to navigate through dark caves where even sensitive vision may not be useful enough."
Pigeon Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
s can hear infrasound. With the average pigeon being able to hear sounds as low as 0.5 Hz, they can detect distant storms, earthquakes and even volcanoes. This also helps them to navigate.


Insects

Greater wax moths (Galleria mellonella) have the highest recorded sound frequency range that has been recorded so far. They can hear frequencies up to 300 kHz. This is likely to help them evade bats.


Fish

Fish have a narrow hearing range compared to most mammals.
Goldfish The goldfish (''Carassius auratus'') is a freshwater fish in the family Cyprinidae of order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the wild have bec ...
and
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
do possess a
Weberian apparatus The Weberian apparatus is an anatomical structure that connects the swim bladder to the auditory system in fishes belonging to the superorder Ostariophysi. When it is fully developed in adult fish, the elements of the apparatus are sometimes co ...
and have a wider hearing range than the
tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae (mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max length: ...
.


Marine mammals

As aquatic environments have very different physical properties than land environments, there are differences in how marine mammals hear compared with land mammals. The differences in
auditory system The auditory system is the sensory system for the sense of hearing. It includes both the sensory organs (the ears) and the auditory parts of the sensory system. System overview The outer ear funnels sound vibrations to the eardrum, increasin ...
s have led to extensive research on aquatic mammals, specifically on dolphins. Researchers customarily divide marine mammals into five hearing groups based on their range of best underwater hearing. (Ketten, 1998): Low-frequency baleen whales like blue whales (7 Hz to 35 kHz); Mid-frequency toothed whales like most dolphins and sperm whales (150 Hz to 160 kHz) ; High-frequency toothed whales like some dolphins and porpoises (275 Hz to 160 kHz); Seals (50 Hz to 86 kHz); Fur seals and sea lions (60 Hz to 39 kHz). The auditory system of a land mammal typically works via the transfer of sound waves through the ear canals. Ear canals in
seals Seals may refer to: * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to impress an emblem, used as a means of a ...
, sea lions, and
walruses The walrus (''Odobenus rosmarus'') is a large flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in the fami ...
are similar to those of land mammals and may function the same way. In whales and dolphins, it is not entirely clear how sound is propagated to the ear, but some studies strongly suggest that sound is channelled to the ear by tissues in the area of the lower jaw. One group of whales, the Odontocetes (toothed whales), use echolocation to determine the position of objects such as prey. The toothed whales are also unusual in that the ears are separated from the skull and placed well apart, which assists them with localizing sounds, an important element for echolocation. Studies have found there to be two different types of cochlea in the dolphin population. Type I has been found in the
Amazon river dolphin The Amazon river dolphin (''Inia geoffrensis''), also known as the boto, bufeo or pink river dolphin, is a species of toothed whale classified in the family Iniidae. Three subspecies are currently recognized: ''I. g. geoffrensis'' (Amazon river ...
and harbour porpoises. These types of dolphin use extremely high frequency signals for echolocation. Harbour porpoises emit sounds at two bands, one at 2 kHz and one above 110 kHz. The cochlea in these dolphins is specialised to accommodate extreme high frequency sounds and is extremely narrow at the base. Type II cochlea are found primarily in offshore and open water species of whales, such as the
bottlenose dolphin Bottlenose dolphins are aquatic mammals in the genus ''Tursiops.'' They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus definitively contains two species: the comm ...
. The sounds produced by bottlenose dolphins are lower in frequency and range typically between 75 and 150,000 Hz. The higher frequencies in this range are also used for echolocation and the lower frequencies are commonly associated with social interaction as the signals travel much farther distances. Marine mammals use vocalisations in many different ways. Dolphins communicate via clicks and whistles, and whales use low-frequency moans or pulse signals. Each signal varies in terms of frequency and different signals are used to communicate different aspects. In dolphins, echolocation is used in order to detect and characterize objects and whistles are used in sociable herds as identification and communication devices.


See also

*
Audiology Audiology (from Latin , "to hear"; and from Greek , ''-logia'') is a branch of science that studies hearing, balance, and related disorders. Audiologists treat those with hearing loss and proactively prevent related damage. By employing vario ...
*
Audiometry Audiometry () is a branch of audiology and the science of measuring hearing acuity for variations in sound intensity and pitch and for tonal purity, involving thresholds and differing frequencies. Typically, audiometric tests determine a subje ...
* The Mosquito * Seismic communication * Minimum audibility curve *
Musical acoustics Musical acoustics or music acoustics is a multidisciplinary field that combines knowledge from physics, psychophysics, organology (classification of the instruments), physiology, music theory, ethnomusicology, signal processing and instrument build ...


Notes


References


Further reading

* * * * * {{Authority control Otology Zoology