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Alderflies are megalopteran insects of the family Sialidae. They are closely related to the
dobsonflies Dobsonflies are a subfamily of insects, Corydalinae, part of the Megalopteran family Corydalidae. The larvae (commonly called hellgrammites) are aquatic, living in streams, and the adults are often found along streams as well. The nine genera o ...
and
fishflies Fishflies are members of the subfamily Chauliodinae, belonging to the megalopteran family Corydalidae. They are most easily distinguished from their closest relatives, dobsonflies, by the jaws (mandibles) and antennae. In contrast to the large ...
as well as to the
prehistoric Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use o ...
Euchauliodidae. All living alderflies – about 66
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
all together – are part of the subfamily Sialinae, which contains between one and seven extant genera according to different scientists' views.


Description

Sialinae have a body length of less than 25 mm (1 inch), long filamentous antennae, and four large dark
wings A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is express ...
of which the
anterior Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
pair is slightly longer than the posterior. They lack ocelli and their fourth tarsal segment is dilated and deeply bilobed. Dead alderfly larvae are used as bait in fishing.fly fishing entomology alderfy entry
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Life cycle

The females lay a vast number of eggs on grass stems near water. When the larvae are born they drop into the water or the ground nearby it and make their way into their new aquatic biome. The larvae are aquatic, active, armed with strong sharp mandibles, and breathe by means of seven pairs of abdominal branchial filaments. When full sized, which takes between one and two years, they leave the water and spend a quiescent pupal stage on the land before metamorphosis into the sexually mature insect. Adult alderflies stay near to the water in which they had lived when they were younger. Once in their adult stage, they tend to live only 2 to 3 weeks, which they spend mainly in reproducing.


Classification

In addition to the seven living genera, there are several genera of fossil alderflies. Family Sialidae *Genus †'' Dobbertinia'' Handlirsch 1920 Green Series, Germany, Early Jurassic (
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarc ...
) *Genus †'' Sharasialis'' Ponomarenko 2012 Shar-Teeg, Mongolia, Late Jurassic ( Tithonian) *Subfamily Sialinae **Genus '' Austrosialis'' Tillyard,
919 __NOTOC__ Year 919 ( CMXIX) was a common year starting on Friday (link will display the full calendar) of the Julian calendar. Events By Place Byzantine Empire * March 25 – Romanos Lekapenos, admiral (''droungarios'') of the Byz ...
/small> ***Species '' Austrosialis ignicollis'' Tillyard,
919 __NOTOC__ Year 919 ( CMXIX) was a common year starting on Friday (link will display the full calendar) of the Julian calendar. Events By Place Byzantine Empire * March 25 – Romanos Lekapenos, admiral (''droungarios'') of the Byz ...
/small> ***Species '' Austrosialis maxmouldsi'' Theischinger, 1983 **Genus †'' Eosialis'' Nel et al., 2002 ***Species †'' Eosialis dorisi'' Nel et al., 2002 **Genus '' Indosialis'' Lestage, 1927 ***Species '' Indosialis bannaensis'' X.-y. Liu et al., 2006 ***Species '' Indosialis beskonakensis'' Nel, 1988 ***Species '' Indosialis indica'' X.-y. Liu et al., 2008 ***Species '' Indosialis minora'' (Banks, 1920) **Genus '' Leptosialis'' Esben-Petersen, 1920 ***Species '' Leptosialis africana Esben-Petersen, 1920'' **Genus '' Nipponosialis'' Kuwayama, 1962 ***Species '' Nipponosialis amamiensis'' Kuwayama, 1964 ***Species '' Nipponosialis jezoensis'' (Okamoto, 1910) ****Subspecies '' Nipponosialis jezoensis jezoensis'' (Okamoto, 1910) ****Subspecies '' Nipponosialis jezoensis kuwayamai'' Hayashi & Suda, 1995 ***Species '' Nipponosialis kumejimae'' (Okamoto, 1910) **Genus †'' Proindosialis'' Nel, 1988 ***Species †'' Proindosialis cantalensis'' Nel, 1988 **Genus '' Protosialis'' van der Weele, 1909 ***Species '' Protosialis afra'' Navás, (1936) ***Species '' Protosialis americana'' (Rambur, 1842) ***Species '' Protosialis australiensis'' (Tillyard, 919) ***Species ''Protosialis australis'' Navás, 1927 ***Species †''Protosialis baltica'' (Wichard, 1997) ***Species ''Protosialis bifasciata'' (Hagen, 1861) ***Species ''Protosialis bimaculata'' Banks, 1920 ***Species ''Protosialis brasiliensis'' Navás, 1936 ***Species †'' Protosialis casca'' (Engel & Grimaldi, 2007) ***Species '' Protosialis chilensis'' (McLachlan, (1871) ***Species '' Protosialis flammata'' Penny, (1982) ***Species '' Protosialis flavicollis'' (Enderlein, 1910) ***Species '' Protosialis hauseri'' Contreras-Ramos et al., 2005 ***Species '' Protosialis madegassa'' Navás, 1927 ***Species '' Protosialis mexicana'' (Banks, 1901) ***Species '' Protosialis minora'' Banks, 1920 ***Species '' Protosialis nubila'' Navás, 1933 ***Species '' Protosialis ranchograndis'' Contreras-Ramos, 2006 ***Species †'' Protosialis voigti'' (Wichard & Engel, 2006) **Genus ''
Sialis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Latreille, 1802 ***Species Approximately 90 ***Species ''
Sialis abchasica ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 ***Species '' Sialis aequalis'' Banks, 1920 ***Species ''
Sialis americana ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' (Rambur, 1842) ***Species '' Sialis annae'' Vshivkova, 1979 ***Species ''
Sialis arvalis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis atra'' Navás, (1928) ***Species ''
Sialis bifida ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Hayashi & Suda, 1997 ***Species ''
Sialis bilineata ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Say, 1823 ***Species '' Sialis bilobata'' Whiting, 1991 ***Species ''
Sialis californica ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Banks, 1920 ***Species ''
Sialis chilensis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' McLachlan, (1871) ***Species '' Sialis concava'' Banks, 1897 ***Species '' Sialis contigua'' Flint, 1964 ***Species ''
Sialis cornuta ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis didyma'' Navás, 1916 ***Species ''
Sialis dorochovae ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 ***Species '' Sialis dorsata'' Say, 1823 ***Species ''
Sialis dreisbachi ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Flint, 1964 ***Species '' Sialis elegans'' X.-y. Liu & D. Yang, 2006 ***Species ''
Sialis flavicollis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Enderlein, 1910 ***Species '' Sialis formosana'' Esben-Petersen, 1913 ***Species '' Sialis frequens'' Okamoto, 1905 ***Species ''
Sialis fuliginosa ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' F. Pictet, 1836 ***Species ''
Sialis fumosa ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Navás, 1915 ***Species '' Sialis glabella'' Ross, 1937 ***Species ''
Sialis gonzalezi ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 ***Species †''
Sialis groehni ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Wichard, 1997 ***Species '' Sialis hamata'' Ross, 1937 ***Species ''
Sialis hasta ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis henanensis'' X.-y. Liu & D. Yang, 2006 ***Species †'' Sialis herrlingi'' Wichard, 2002 ***Species '' Sialis imbecilla'' Say, 1823 ***Species ''
Sialis immarginata ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Say, 1823 ***Species ''
Sialis infumata ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Newman, 1838 ***Species '' Sialis iola'' Ross, 1937 ***Species ''
Sialis itasca ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis japonica'' van der Weele, 1909 ***Species '' Sialis jianfengensis'' D. Yang et al., 2002 ***Species ''
Sialis joppa ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species ''
Sialis klingstedti ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 ***Species '' Sialis kunmingensis'' X.-y. Liu & D. Yang, 2006 ***Species ''
Sialis levanidovae ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1980 ***Species ''
Sialis longidens ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Klingstedt, (1932) ***Species ''
Sialis lutaria ''Sialis lutaria'', common name alderfly, is a species of alderfly belonging to the order Megaloptera family Sialidae. Distribution This species is mainly present in Austria, Belgium, United Kingdom, Czech Republic, France, Germany, Italy, t ...
'' (Linnaeus, 1758) ***Species ''
Sialis martynovae ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1980 ***Species '' Sialis melania'' Nakahara, 1915 ****Subspecies '' Sialis melania kyushuensis'' Hayashi & Suda, 1995 ****Subspecies '' Sialis melania melania'' Nakahara, 1915 ****Subspecies '' Sialis melania tohokuensis'' Hayashi & Suda, 1995 ****Subspecies '' Sialis melania toyamaensis'' Hayashi & Suda, 1995 ***Species '' Sialis mohri'' Ross, 1937 ***Species ''
Sialis morio ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Klingstedt, (1933) ***Species '' Sialis morrisoni'' K. Davis, 1903 ***Species '' Sialis muratensis'' Nel, 1988 ***Species ''
Sialis nevadensis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' K. Davis, 1903 ***Species ''
Sialis nigripes ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' E. Pictet, 1865 ***Species '' Sialis nina'' Townsend, 1939 ***Species '' Sialis occidens'' Ross, 1937 ***Species '' Sialis rotunda'' Banks, 1920 ***Species '' Sialis sibirica'' McLachlan, 1872 ***Species ''
Sialis sinensis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Banks, (1940) ***Species '' Sialis sordida'' Klingstedt, (1933) ***Species ''
Sialis spangleri ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Flint, 1964 ***Species '' Sialis strausi'' Illies, 1967 ***Species '' Sialis vagans'' Ross, 1937 ***Species ''
Sialis vanderweelei ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' U. Aspöck & H. Aspöck, 1983 ***Species ''
Sialis velata ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis versicoloris'' X.-y. Liu & D. Yang, 2006 ***Species ''
Sialis yamatoensis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Hayashi & Suda, 1995 ***Species ''
Sialis zhiltzovae ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 **Genus '' Stenosialis'' Tillyard, (1919) ***Species '' Stenosialis australiensis'' Tillyard, (1919) ***Species '' Stenosialis hollowayi'' Theischinger, 1983 ''Sialis lutaria'' is the commonest alderfly in the United Kingdom (2006)
Alderfly - ''Sialis lutaria''
Retrieved 28 June 2006.
and across much of Europe.


References


External links


BugGuide


{{Authority control Megaloptera Aquatic insects