The Terek sandpiper (''Xenus cinereus'') is a small migratory

Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

wader 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

and is the only member of the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

''Xenus''. It is named after the Terek River which flows into the west of the

Caspian Sea The Caspian Sea is the world's largest inland body of water, described as the List of lakes by area, world's largest lake and usually referred to as a full-fledged sea. An endorheic basin, it lies between Europe and Asia: east of the Caucasus, ...

, as it was first observed around this area.

Taxonomy

The Terek sandpiper was formally described and illustrated in 1775 by the German naturalist

Johann Anton Güldenstädt Johann Anton Güldenstädt (26 April 1745 in Riga, Latvia – 23 March 1781 in St. Petersburg, Russia) was a Baltic German natural history, naturalist and explorer in Russian service. Güldenstädt lost both his parents early, and from 1763 on ...

under the

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...

''Scolopax cinerea''. He reported that he had seen pairs breeding at the mouth of the Terek River where it flows into the

. It is now the only species placed in the

''Xenus'' that was introduced in 1829 by the German naturalist

Johann Jakob Kaup Johann Jakob von Kaup (10 April 1803 – 4 July 1873) was a German naturalist. A proponent of natural philosophy, he believed in an innate mathematical order in nature and he attempted biological classifications based on the Quinarian system. Kaup ...

. The genus name ''Xenus'' is from

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

''xenos'' meaning "stranger"; the specific epithet ''cinereus'' is

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

for "ash-grey" from ''cinis, cineris'', "ashes". The species is considered to be

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

, no

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

are recognised. Among the

Scolopacidae Scolopacidae is a large family of shorebirds, or waders, which mainly includes many species known as sandpipers, but also others such as woodcocks, curlews and snipes. Most of these species eat small invertebrates picked out of the mud or soil ...

, ''Xenus'' is part of the shank- tattler-

phalarope A phalarope is any of three living species of slender-necked shorebirds in the genus ''Phalaropus'' of the bird family Scolopacidae. Phalaropes are close relatives of the shanks and tattlers, the '' Actitis'' and Terek sandpipers, and also ...

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

and less closely related to the calidrid sandpipers. Based on the degree of

DNA sequence A nucleic acid sequence is a succession of bases within the nucleotides forming alleles within a DNA (using GACT) or RNA (GACU) molecule. This succession is denoted by a series of a set of five different letters that indicate the order of the nu ...

divergence and putative shank and phalarope

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s from around the

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

boundary some 23–22

million years ago Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used w ...

, the Terek sandpiper presumably diverged from their relatives in the

Late Oligocene The Chattian is, in the geologic timescale The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the pro ...

. Given the numerous basal fossils of the group found in

Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...

it is likely that the Terek sandpiper lineage originated there, possibly by being isolated as the remains of the Turgai Sea dried up, which happened just around this time.

Description

Slightly larger than the common sandpiper at length, its long upcurved bill – somewhat reminiscent of an avocet's, but not as strongly curved – makes it very distinctive. As the scientific specific name implies, this wader has a grey back, face and breast in all

plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...

s; a white

supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...

may appear more or less distinct. The belly is whitish and the feet yellow; the bill has a yellowish base, with the rest being black. The call is a high whistle.

Distribution and ecology

This bird breeds near water in the

taiga Taiga or tayga ( ; , ), also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches. The taiga, or boreal forest, is the world's largest land biome. In North A ...

from

Finland Finland, officially the Republic of Finland, is a Nordic country in Northern Europe. It borders Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bothnia to the west and the Gulf of Finland to the south, ...

through northern

Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...

to the

Kolyma River The Kolyma (, ; ) is a river in northeastern Siberia, whose basin covers parts of the Sakha Republic, Chukotka Autonomous Okrug, and Magadan Oblast of Russia. The Kolyma is frozen to depths of several metres for about 250 days each year, b ...

, and

migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...

south in winter to tropical coasts in east Africa, south Asia and

Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...

, usually preferring muddy areas. It is a rare vagrant in western Europe, and particularly in autumn it is sometimes seen passing through the

Marianas The Mariana Islands ( ; ), also simply the Marianas, are a crescent-shaped archipelago comprising the summits of fifteen longitudinally oriented, mostly Volcano#Dormant and reactivated, dormant volcanic mountains in the northwestern Pacific Ocean ...

on migration; on

Palau Palau, officially the Republic of Palau, is an island country in the Micronesia subregion of Oceania in the western Pacific Ocean. The Republic of Palau consists of approximately 340 islands and is the western part of the Caroline Islands ...

, further off its usual migration route, it is decidedly uncommon on the other hand. Almost annually and apparently more and more often in recent times, a few birds stray to

Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...

and the Aleutian and

Pribilof Islands The Pribilof Islands (formerly the Northern Fur Seal Islands; , ) are a group of four volcanic islands off the coast of mainland Alaska, in the Bering Sea, about north of Unalaska and 200 miles (320 km) southwest of Cape Newenham. The ...

. Every few years, individual vagrants are recorded in the

Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeog ...

, where they arrive either as migrating birds from Africa, or as North American strays accompanying local waders south for winter. Such vagrants have been recorded as far south as

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

. The overall genetic variation in Terek sandpipers across their range is low, with some evidence of contractions followed by expansion. Although the geographically isolated Dnieper River population in Eastern Europe does show significant genetic differentiation. It feeds in a distinctive and very active way, chasing insects and other mobile prey, and sometimes then running to the water's edge to wash its catch. It lays three or four eggs in a lined ground scrape. The Terek sandpiper likes to associate with

ruddy turnstone The ruddy turnstone (''Arenaria interpres'') is a small Cosmopolitan distribution, cosmopolitan wader, wading bird, one of two species of turnstone in the genus ''Arenaria''. It is now classified in the sandpiper family Scolopacidae but was form ...

s (''Arenaria interpres''), smallish calidrids, and ''

Charadrius ''Charadrius'' is a genus of plovers, a group of wading birds. The genus name ''Charadrius'' is a Late Latin word for a yellowish bird mentioned in the fourth-century Vulgate. They are found throughout the world. Many ''Charadrius'' species are ...

'' (but maybe not '' Pluvialis'')

plover Plovers ( , ) are members of a widely distributed group of wader, wading birds of subfamily Charadriinae. The term "plover" applies to all the members of the subfamily, though only about half of them include it in their name. Species lis ...

s; a vagrant bird at

Paraty Paraty (or Parati, ) is a preserved Portuguese colonial (1500–1822) and Brazilian Imperial (1822–1889) municipality with a population of about 43,000. The name "Paraty" originates from the local Guaianá Indians' indigenous Tupi language, ...

(

Rio de Janeiro state Rio de Janeiro () is one of the 27 federative units of Brazil. It has the second largest economy of Brazil, with the largest being that of the state of São Paulo. The state, which has 8.2% of the Brazilian population, is responsible for 9.2% of ...

) was noted to pair up with a

spotted sandpiper The spotted sandpiper (''Actitis macularius'') is a small shorebird. Together with its sister species the common sandpiper (''A. hypoleucos''), it makes up the genus ''Actitis''. They replace each other geographically; stray birds may Hybridisati ...

(''Actitis macularius''). This is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' (

AEWA The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA), is an independent international treaty developed under the auspices of the United Nations Environment Programme's Convent ...

) applies. Widespread and often quite commonly seen, the Terek sandpiper is not considered a threatened species by the

IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

Gallery

File:Terek Sandpiper, Little Stint, Marsh Sandpiper, C Redshank W2 IMG 6891.jpg, Terek sandpiper ''Xenus cinereus'',

little stint The little stint (''Calidris minuta'' or ''Erolia minuta'') is a very small wader. It breeds in arctic Europe and Asia, and is a long-distance migrant, wintering south to Africa and south Asia. It occasionally is a vagrant to North America an ...

''Calidris minuta'' and

marsh sandpiper The marsh sandpiper (''Tringa stagnatilis'') is a small wader. It is a rather small Tringa, shank, and breeds in open grassy steppe and taiga wetlands from easternmost Europe to the Russian Far East. The genus name ''Tringa'' is the Neo-Latin name ...

''Tringa stagnatilis'' in Krishna Wildlife Sanctuary,

Andhra Pradesh Andhra Pradesh (ISO 15919, ISO: , , AP) is a States and union territories of India, state on the East Coast of India, east coast of southern India. It is the List of states and union territories of India by area, seventh-largest state and th ...

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

. File:Xenus cinereus eating ragworm.JPG, Eating ragworm in

Shōnai River The is a Class 1 river flowing through Gifu and Aichi prefectures in Japan. In Gifu Prefecture, it is also referred to as the Toki River (土岐川 ''Toki-gawa''); around the city of Kasugai in Aichi Prefecture, it is referred to as the Tam ...

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

. File:Xenus cinereus MWNH 0248.JPG, Egg, Collection

Museum Wiesbaden The Museum Wiesbaden is a two-branch museum of Art museum, art and Natural history museum, natural history in the Hesse, Hessian capital of Wiesbaden, Germany. It is one of the three Hessian State museums, in addition to the museums in Hessian ...

File:Terek Sandpiper (Xenus cinereus) (8079446567).jpg, In flight File:Terek Sandpiper AMSM5480 TSAN.jpg, In Narara,

Jamnagar Jamnagar () is a city and the headquarters of Jamnagar district in the Indian state of Gujarat. The city lies just to the south of the Gulf of Kutch, some west of the state capital, Gandhinagar. The city was the capital of Nawanagar State, Na ...

File:Terek Sandpiper AMSM5481.jpg, In Narara,

File:Xenus cinereus (Alnus).jpg, In

Yunlin County Yunlin is a Counties of Taiwan, county in Regions of Taiwan, western Taiwan. Yunlin County borders the Taiwan Strait to the west, Nantou County to the east, Changhua County to the north at the Zhuoshui River, and Chiayi County to the south at th ...

, Taiwan.

References

External links

Terek sandpiper species text in The Atlas of Southern African Birds
* * * * * * * {{Taxonbar, from=Q18856

Terek sandpiper The Terek sandpiper (''Xenus cinereus'') is a small migratory Palearctic wader 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly foun ...

Birds of Russia Birds of Central Asia