''Widdringtonia'' is a genus of coniferous

tree In botany, a tree is a perennial plant with an elongated Plant stem, stem, or trunk (botany), trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondar ...

s in the

Cupressaceae Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioeci ...

( cypress family). The name was Austrian botanist

Stephan Endlicher Stephan Ladislaus Endlicher also known as Endlicher István László (24 June 1804, Bratislava (Pozsony) – 28 March 1849, Vienna) was an Austrian botanist, numismatist and Sinologist. He was a director of the Botanical Garden of Vienna. B ...

's way of honouring an early expert on the coniferous forests of Spain, Capt. Samuel Edward Widdrington (1787–1856). There are four species, all native to southern

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, where they are known as cedars or African cypresses.

Description

This genus contains large evergreen

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

s and

s, reaching 5–20 m tall (to 40 m in ''W. whytei''). Juveniles have needle-like leaves that are arranged in spirals. The scale-like leaves in adults are arranged in decussate opposite pairs in four rows along the twigs. The plants are dioecious. The small male cones grow at the ends of twigs. The scales have no stalks. They grow into an upward, decreasing beak with two to six pollen sacs at the base of the cone. The female cones are small and stalkless and grow in short spikes on the branches. The scales grow in two opposite rows that spread at the base during pollination. They then close into a corky, leathery cone with five or more ovules at the base of each scale. The cones become woody as they mature. The cones mostly remain closed on the trees for many years, opening only after being scorched by a

wildfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identi ...

; this then releases the

seed A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, includ ...

s to grow on the newly cleared burnt ground. They open at four very thick valves that correspond to the four scales. Each cone produces few seeds. They are wing with hard test and two cotyledons. In ''W. whytei'' the cones open soon after maturity to shed the seed without fire; this species is more sensitive to fire and only grows in moister situations where it is protected from fire. The best adapted to fire is ''W. nodiflora'', which has the ability to re-grow from the roots, as well as by seed.

Species

One species is widespread in southern Africa, while the other three have restricted ranges, often occurring with or near to the widespread species. The closest relatives of ''Widdringtonia'' are '' Callitris'' and ''

Actinostrobus ''Actinostrobus'' is a genus of coniferous trees in the Cupressaceae (cypress family). Common names include cypress, sandplain-cypress and cypress-pine, the last of these shared by the closely related genus ''Callitris''. Species There are three ...

'' from Australia, which differ in their cones and leaves being in whorls of three, not opposite pairs.

Uses

The wood is light, soft and aromatic. It can be easily split and resists decay. It is used to make furniture, indoor and outdoor panelling, and fence posts. That of ''W. whytei'' was particularly valuable as it was available in large sizes, but this species is now endangered and no longer cut to any extent.

References and external links

* Pauw, C. A. & Linder, H. P. 1997. ''Widdringtonia'' systematics, ecology and conservation status. ''Bot. J. Linn. Soc.'' 123: 297–319.
Recommended English names for trees of Southern Africa
(archive)

{{Taxonbar, from=Q117681 Afromontane flora Conifer genera Trees of South Africa Taxa named by Stephan Endlicher