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In botany, a whorl or verticil is an arrangement of leaves,
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...
s, petals,
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fila ...
s, or carpels that radiate from a single point and surround or wrap around the stem or stalk. A leaf whorl consists of at least three elements; a pair of opposite leaves is not called a whorl. For leaves to grow in whorls is fairly rare except in plant species with very short
internodes A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrien ...
and some other genera ( Galium,
Nerium ''Nerium oleander'' ( ), most commonly known as oleander or nerium, is a shrub or small tree cultivated worldwide in temperate and subtropical areas as an ornamental and landscaping plant. It is the only species currently classified in the ge ...
, Elodea etc.). Leaf whorls occur in some trees such as '' Brabejum stellatifolium'' and other species in the family Proteaceae (e.g., in the genus '' Banksia''). In plants such as these, crowded internodes within the leaf whorls alternate with long internodes between the whorls. The morphology of most
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s (called
cyclic flower A cyclic flower is a flower type formed out of a series of whorls; sets of identical organs attached around the axis at the same point. Most flowers consist of a single whorl of sepals termed a calyx; a single whorl of petals termed a corolla; one ...
s) is based on four types of whorls: # The
calyx Calyx or calyce (plural "calyces"), from the Latin ''calix'' which itself comes from the Ancient Greek ''κάλυξ'' (''kálux'') meaning "husk" or "pod", may refer to: Biology * Calyx (anatomy), collective name for several cup-like structures ...
: zero or more whorls of sepals at the base # The
corolla Corolla may refer to: *Corolla (botany), the petals of a flower, considered as a unit *Toyota Corolla, an automobile model name *Corolla (headgear) A ''corolla'' is an ancient headdress in the form of a small circlet or crown.androecium The stamen ( plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fil ...
: zero or more whorls of stamens, each comprising a filament and an anther # The
gynoecium Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pist ...
: zero or more whorls of carpels, each consisting of an
ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...
, a style, and a stigma A flower lacking any of these floral structures is said to be ''incomplete'' or ''imperfect''. Not all flowers consist of whorls since the parts may instead be spirally arranged, as in the family Magnoliaceae.


See also

*
Cyclic flower A cyclic flower is a flower type formed out of a series of whorls; sets of identical organs attached around the axis at the same point. Most flowers consist of a single whorl of sepals termed a calyx; a single whorl of petals termed a corolla; one ...
* Merosity * Phyllotaxis


References

Plant morphology {{botany-stub