Wastebasket Genus
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Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some
taxonomist In biology, taxonomy () is the science, scientific study of naming, defining (Circumscription (taxonomy), circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxon, taxa (si ...
s to refer to a
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
that has the purpose of classifying organisms that do not fit anywhere else. They are typically defined by either their designated members' often superficial similarity to each other, or their ''lack'' of one or more distinct character states or by their ''not'' belonging to one or more other taxa. Wastebasket taxa are by definition either
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
or
polyphyletic A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...
, and are therefore not considered valid taxa under strict
cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...
rules of taxonomy. The name of a wastebasket taxon may in some cases be retained as the designation of an
evolutionary grade A grade is a taxon united by a level of morphological or physiological complexity. The term was coined by British biologist Julian Huxley, to contrast with clade, a strictly phylogenetic unit. Phylogenetics The concept of evolutionary grades ...
, however.


Examples

There are many examples of paraphyletic groups, but true "wastebasket" taxa are those that are known not to, and perhaps not intended to, represent natural groups, but are nevertheless used as convenient groups of organisms. The acritarchs are perhaps the most famous example. Wastebasket taxa are often old (and perhaps not described with the systematic rigour and precision that is possible in the light of accumulated knowledge of diversity) and populous. * The Flacourtiaceae, a now-defunct family of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s – the Angiosperm Phylogeny Group has placed its
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
s and genera in various other families, especially the Achariaceae and
Salicaceae The Salicaceae are the willow family of flowering plants. The traditional family (Salicaceae ''sensu stricto'') includes the willows, poplars. Genetic studies summarized by the Angiosperm Phylogeny Group (APG) have greatly expanded the circumsc ...
. * The obsolete kingdom
Protista A protist ( ) or protoctist is any Eukaryote, eukaryotic organism that is not an animal, Embryophyte, land plant, or fungus. Protists do not form a Clade, natural group, or clade, but are a Paraphyly, paraphyletic grouping of all descendants o ...
is composed of all
eukaryote The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms ...
s that are not
animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s,
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s or
fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
, leaving to the protists all single-celled eukaryotes. * The Tricholomataceae is a fungal group, at one point composed of the white-, yellow-, or pink-spored genera in the Agaricales not already classified as belonging to the Amanitaceae, Lepiotaceae, Hygrophoraceae, Pluteaceae, or Entolomataceae. * Carnosauria and Thecodontia are fossil groups, banded together back when the limited fossil record did not allow for a more detailed scheme. * Condylarthra is an artificial clade into which ungulate mammals not clearly within Perissodactyla or
Cetartiodactyla Artiodactyls are placental mammals belonging to the order (biology), order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof ...
were traditionally shoved. Many of these groups, like Meridiungulata or '' Protungulatum'', may not represent laurasitherian mammals, while others like phenacodontids have been clearly established as early odd-toed ungulates. * The order
Insectivora The Order (biology), order Insectivora (from Latin ''insectum'' "insect" and ''vorare'' "to eat") is a now-abandoned biological grouping within the class of mammals. Some species have now been moved out, leaving the remaining ones in the order ...
has traditionally been used as a dumping ground for placental insectivorous mammals (and similar forms such as colugos), usually aligned with
carnivoran Carnivora ( ) is an order of placental mammals specialized primarily in eating flesh, whose members are formally referred to as carnivorans. The order Carnivora is the sixth largest order of mammals, comprising at least 279 species. Carnivor ...
s,
ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
s and bats. While the core components ( moles,
shrew Shrews ( family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to dif ...
s,
hedgehog A hedgehog is a spiny mammal of the subfamily Erinaceinae, in the eulipotyphlan family Erinaceidae. There are 17 species of hedgehog in five genera found throughout parts of Europe, Asia, and Africa, and in New Zealand by introduction. The ...
s and their close relations) do in fact form a consistent clade, Eulipotyphla, that is part of
Laurasiatheria Laurasiatheria (; "Laurasian beasts") is a superorder of Placentalia, placental mammals that groups together true insectivores (eulipotyphlans), bats (chiropterans), carnivorans, pangolins (Pholidota, pholidotes), even-toed ungulates (Artiodacty ...
with the aforementioned clades, other mammals historically placed in the order have been found to belong to other branches of the placental tree:
tree shrew The treeshrews (also called tree shrews or banxrings) are small mammals native to the tropical forests of South and Southeast Asia. They make up the entire Order (biology), order Scandentia (from Latin ''scandere'', "to climb"), which split into ...
s and colugos are euarchontans related to
Primates Primates is an order of mammals, which is further divided into the strepsirrhines, which include lemurs, galagos, and lorisids; and the haplorhines, which include tarsiers and simians ( monkeys and apes). Primates arose 74–63  ...
and sometimes grouped in Sundatheria, while tenrecs, golden moles and elephant shrews are all afrotheres, probably forming the clade Afroinsectiphilia. Both of these clades have at times been accused of being wastebasket taxa themselves, grouping superficially similar animals in Euarchonta and Afrotheria, respectively, but they have been more strongly supported by genetic studies. * Vermes is an obsolete taxon of worm-like animals. It was a catch-all term used by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
and
Jean-Baptiste Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biologi ...
for non-arthropod invertebrate animals. *The genus '' Mamenchisaurus'' is sometimes considered a wastebasket taxon for large, long-necked
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s.


Wastebasket taxa in science

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
groups that are poorly known due to fragmentary remains are sometimes grouped together on gross morphology or
stratigraphy Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithost ...
, only later to be found to be wastebasket taxa, such as the crocodile-like
Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
group Rauisuchia. One of the roles of taxonomists is to identify wastebasket taxa and reclassify the content into more natural units. Sometimes, during taxonomic revisions, a wastebasket taxon can be salvaged after doing thorough research on its members, and then imposing tighter restrictions on what continues to be included. Such techniques "saved" Carnosauria and ''
Megalosaurus ''Megalosaurus'' (meaning "great lizard", from Ancient Greek, Greek , ', meaning 'big', 'tall' or 'great' and , ', meaning 'lizard') is an extinct genus of large carnivorous theropod dinosaurs of the Middle Jurassic Epoch (Bathonian stage, 166 ...
''. Other times, the taxonomic name contains too much unrelated "baggage" to be successfully salvaged. As such, it is usually dumped in favour of a new, more restrictive name (for example, Rhynchocephalia), or abandoned altogether (for example, '' Simia'').


Related concepts

A related concept is that of
form taxon Form classification is the classification of organisms based on their morphology (biology), morphology, which does not necessarily reflect their biological relationships. Form classification, generally restricted to palaeontology, reflects uncer ...
, "wastebasket" groupings that are united by gross morphology. This is often result of a common mode of life, often one that is generalist, leading to generally similar body shapes by
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
. The term wastebasket taxon is sometimes employed in a derogatory fashion to refer to an
evolutionary grade A grade is a taxon united by a level of morphological or physiological complexity. The term was coined by British biologist Julian Huxley, to contrast with clade, a strictly phylogenetic unit. Phylogenetics The concept of evolutionary grades ...
taxon.


See also

*
Lazarus taxon In paleontology, a Lazarus taxon (plural ''taxa'') is a taxon that disappears for one or more periods from the fossil record, only to appear again either in later fossil records, or as actual living organisms, and often in isolated, obscure, ...
* Elvis taxon * ''
Incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
'' *
Glossary of scientific naming This is a list of terms and symbols used in scientific names for organisms, and in describing the names. For proper parts of the names themselves, see List of Latin and Greek words commonly used in systematic names. Many of the abbreviations are ...
* Not Otherwise Specified


References

{{Reflist, 2 Obsolete taxa Taxonomy (biology) Paraphyletic groups Polyphyletic groups