''Variola'', the lyretails, is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...

grouper Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...

s from the subfamily

Epinephelinae Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...

, part of the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some ca ...

, which also includes the

anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name Anthi ...

and sea basses. They are found in the tropical Indo-Pacific and their distribution extends from the

Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; ...

to South Africa across the Indian Ocean and east to the islands of the central Pacific.

Characteristics

The lyretails have an oblong shaped body which has a depth that is less than the length of the head and which has a standard length that is 2.8 to 3.2 times its depth. They have a slightly convex dorsal profile of the head and a snout which is longer than the diameter of the eye. Both species have a rounded preopercle with a finely serrated margin and a fleshy lower margin. There are three flat spines on the gill cover which has a straight upper margin. Both the upper and lower jaws have a pair of large canine teeth at the front and there are also 1 to 3 large canines in the middle of the lower jaw There are teeth on the roof of the mouth. The

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...

contains 9 spines and 13 to 15 soft rays and it has its origin above the posterior end of the gill cover. The membranes between the spines of the dorsal fin are not or are only slightly incised. The anal fin has 3 separated spines and 8 soft rays. The

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

is crescent shaped with elongated lobes.

Habitat and biology

''Variola'' species occur at depths of over coral reefs where they are normally observed seen swimming some distance above the reef and they appear to prefer areas with clear water such as islands and offshore reefs. They are largely

piscivorous A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evoluti ...

Species

The two known species are:

References

{{Taxonbar, from=Q137222 Epinephelini