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''Ulemosaurus'' is an extinct genus of
dinocephalia Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephal ...
n
therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...
s that lived 265 to 260 million years ago, at Isheevo in
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
n
Tatarstan The Republic of Tatarstan (russian: Республика Татарстан, Respublika Tatarstan, p=rʲɪsˈpublʲɪkə tətɐrˈstan; tt-Cyrl, Татарстан Республикасы), or simply Tatarstan (russian: Татарстан, tt ...
. It was a
tapinocephalid Tapinocephalidae was an advanced family of tapinocephalians. It is defined as the clade containing ''Ulemosaurus'', '' Tapinocaninus'', and the Tapinocephalinae. They are known from both Russia and South Africa. In all probability, the Tapinoceph ...
, a group of bulky herbivores which flourished in the
Middle Permian The Guadalupian is the second and middle series/ epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± ...
. ''Ulemosaurus'' and other
tapinocephalia The Tapinocephalia are one of the major groups of dinocephalian therapsids and the major herbivorous group. Tapinocephalia has been found to consist of three clades: Styracocephalidae, Titanosuchidae, and the very successful Tapinocephalidae. No ...
ns disappeared at the end of the
Middle Permian The Guadalupian is the second and middle series/ epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± ...
.


Description

Only several partial skeletons and skulls have been found. The skull bones are extremely dense: about at its thickest. This thickening is possibly related to head-butting behavior, as some researchers suggest. The species is considered a
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...
, but because the
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
is heavily constructed some
palaeontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of foss ...
s consider it a
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other s ...
, with the species being able to use muscle power to cut prey up with its
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s.


Classification

''Ulemosaurus'' is a large ''
Moschops ''Moschops'' (Greek for "calf face") is an extinct genus of therapsids that lived in the Guadalupian epoch, around 265–260 million years ago. They were heavily built plant eaters, and they may have lived partly in water, as hippopotamuses do. ...
''-like form from Russia; it is probably similar enough to be included as a separate species of ''Moschops''. Despite its advanced characteristics, it lived slightly before the Karoo forms, showing that the Moschopines, and indeed the Tapinocephalidae in general, had already attained their acme by early
Capitanian In the geologic timescale, the Capitanian is an age or stage of the Permian. It is also the uppermost or latest of three subdivisions of the Guadalupian Epoch or Series. The Capitanian lasted between and million years ago. It was preceded by ...
time.


See also

*
List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera tha ...


References


Further reading

* *Riabinin, A. N., 1938. Vertebrate fauna from the Upper Permian deposits of the Sviaga basin: 1. A new Dinocephalian, Ulemosaurus sviagensi n. gen. n. sp: Ezheg. Muz. Akad. F. N. Chernysheva, v. 1, p. 4–40.


External links


Taxonomy
{{Taxonbar, from=Q241603 Tapinocephalians Prehistoric therapsid genera Guadalupian synapsids Extinct animals of Russia Fossil taxa described in 1938 Capitanian genus first appearances Capitanian genus extinctions it:Ulemosaurus