The tympanum is an external hearing structure in animals such as

mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s, some

reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...

s, some

amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...

s and some

insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...

s. Using sound, vertebrates and many insects are capable of sensing their prey, identifying and locating their predators, warning other individuals, and locating potential mates and rivals by hearing the intentional or unintentional sounds they make. In general, any animal that reacts to sounds or communicates by means of sound, needs to have an auditory mechanism. This typically consists of a membrane capable of

vibration Vibration () is a mechanical phenomenon whereby oscillations occur about an equilibrium point. Vibration may be deterministic if the oscillations can be characterised precisely (e.g. the periodic motion of a pendulum), or random if the os ...

known as the tympanum, an air-filled chamber and sensory organs to detect the auditory stimuli.

Insects

Amphibia

Anura

frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...

s and toads, the tympanum is a large external oval shape membrane made up of nonglandular skin. It is located just behind the eye. It does not process

sound wave In physics, sound is a vibration that propagates as an acoustic wave through a transmission medium such as a gas, liquid or solid. In human physiology and psychology, sound is the ''reception'' of such waves and their ''perception'' by the ...

s; it simply transmits them to the inner parts of the amphibian's ear, which is protected from the entry of water and other foreign objects. A frog's ear drum works in very much the same way as does a human eardrum. It is a membrane that is stretched across a ring of cartilage like a snare drum that vibrates. Crossing the middle ear chamber there is an

ossicle The ossicles (also called auditory ossicles) are three irregular bones in the middle ear of humans and other mammals, and are among the smallest bones in the human body. Although the term "ossicle" literally means "tiny bone" (from Latin ''ossicu ...

called the columella that is connected to the tympanum, and another ossicle, the operculum, that connects this to the oval membrane. This separates the middle ear from the

inner ear The inner ear (internal ear, auris interna) is the innermost part of the vertebrate ear. In vertebrates, the inner ear is mainly responsible for sound detection and balance. In mammals, it consists of the bony labyrinth, a hollow cavity in the ...

; and its movements are reflected in vibrations in the fluid in the inner ear; these vibrations cause microscopic hairs to move, which send signals to the frog's brain. A frog's lungs are also involved in the reception of sound, although they are less sensitive than the frog's ear drums.

Insects

Amphibia

Anura

Tetrapoda

Mammals

See also

References

Further reading