Throat clamp is a method of subduing often seen in predatory

felids Felidae () is the family of mammals in the order Carnivora colloquially referred to as cats, and constitutes a clade. A member of this family is also called a felid (). The term "cat" refers both to felids in general and specifically to th ...

and occasionally

canids Canidae (; from Latin, ''canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found within th ...

and

hyaenids Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the clas ...

. It involves the predator using its

jaw The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth and serv ...

to grasp the throat of the prey and clamp tight so that the prey's

windpipe The trachea, also known as the windpipe, is a cartilaginous tube that connects the larynx to the bronchi of the lungs, allowing the passage of air, and so is present in almost all air-breathing animals with lungs. The trachea extends from the l ...

is either crushed or blocked, causing

asphyxiation Asphyxia or asphyxiation is a condition of deficient supply of oxygen to the body which arises from abnormal breathing. Asphyxia causes generalized hypoxia, which affects primarily the tissues and organs. There are many circumstances that can ...

. Cats use this to kill prey while dogs and hyenas use this to weaken the prey before eating it, generally alive. It's more often used than the muzzle clamp and is generally safer, though slower. It is usually most effective when positioned as near to the

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...

as the carnivore can get. Between the larynx and the jaw, the windpipe is surrounded with less

cartilage Cartilage is a resilient and smooth type of connective tissue. In tetrapods, it covers and protects the ends of long bones at the joints as articular cartilage, and is a structural component of many body parts including the rib cage, the neck an ...

and is more malleable, while lower down, near the

chest The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the crea ...

, the passageway would be increasingly harder to collapse, so the throat clamp is usually positioned high up on the animal's neck.

Usage

If the prey is on the ground and pinned, most predators position themselves behind the animal to use their body's weight to help control the movements of the prey and reach around to grab the throat, usually twisting the head around, aiding in both their grasp and the blocking of the windpipe. If the prey is standing, a solitary predator can use a throat clamp usually only if the prey is small or in

shock Shock may refer to: Common uses Collective noun *Shock, a historic commercial term for a group of 60, see English numerals#Special names * Stook, or shock of grain, stacked sheaves Healthcare * Shock (circulatory), circulatory medical emerge ...

. A strong, large animal can easily kick and injure the predator from this angle. A social predator can use the standing throat clamp much more easily because there can be other individuals on the back of the prey, stopping it from kicking, which could lead to collapse, or using a throat clamp periodically to tire the animal out. This latter method usually involves large, patient carnivores such as the

gray wolf The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...

(''Canis lupus'') and

African wild dog The African wild dog (''Lycaon pictus''), also called the painted dog or Cape hunting dog, is a wild canine which is a native species to sub-Saharan Africa. It is the largest wild canine in Africa, and the only extant member of the genus '' Lyc ...

(''Lycaon pictus''). They surround the prey in relatively large numbers, individuals taking turns to jump in and grab the prey by the flanks to try and pull it down, while the prey forcing them off grows exhausted, and in the meantime, more experienced individuals secure a throat clamp on the distracted animal and hold on as long as possible to the struggling prey. The temporary throat clamp can make the prey dizzy and uncoordinated and even force it to go into shock if the chase and attack wasn't enough, aiding in the predator's attempts to pull it to the ground. Agile predators such a lions have been observed to perch on the upper back and neck of large cape buffalo and wrap around to secure a partial throat clamp.

Anatomy

Strong canines are generally required for this move: canines that are too short won't penetrate deeply enough and struggling prey can tear free, while canines that are too long or weak can be broken in the struggle with large prey. This fact created a controversy that is still generally unresolved as to the killing tactics of the extinct carnivores

machairodont Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae (true cats). They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million unt ...

s, or sabre-toothed cats. Strong jaws are often needed to compress the windpipe far enough. Most often, the canines pierce the prey behind the windpipe so that it is not torn, but compressed between premolars of upper and lower jaws. It can be compared to the

bit The bit is the most basic unit of information in computing and digital communications. The name is a portmanteau of binary digit. The bit represents a logical state with one of two possible values. These values are most commonly represented ...

of a horse's bridle in placement. With the mouth completely enclosing the windpipe, it is, if the predator does not relax its grip, very difficult to break free from.

Alternative forms

Occasionally, and as suggested for the killing method of machairodonts, the canines actually pierce the windpipe and the surrounding blood vessels so that the tear in

arteries An artery (plural arteries) () is a blood vessel in humans and most animals that takes blood away from the heart to one or more parts of the body (tissues, lungs, brain etc.). Most arteries carry oxygenated blood; the two exceptions are the p ...

and adjacent windpipe lead to a flow of blood into the windpipe and down into the lungs where the animal essentially drowns in its own blood. This method has not been used regularly by any modern carnivores and is, due to the bloody nature, not suitable for an area with high competition due to the smell attracting other predators. It can be argued that this is not a true throat clamp, but another specialized type of killing.

References

{{reflist Predation