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The mollymawks are a group of medium-sized
albatross Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Pac ...
es that form the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial no ...
''Thalassarche''. The name has sometimes been used for the genus ''
Phoebetria The sooty albatrosses are small albatrosses from the genus ''Phoebetria''. There are two species, the sooty albatross, ''Phoebetria fusca'', and the light-mantled albatross, ''Phoebetria palpebrata''. Systematics The sooties have long been c ...
'' as well, but these are usually called sooty albatrosses. They are restricted to the Southern Hemisphere, where they are the most common of the albatrosses. They were long considered to be in the same genus as the
great albatross The great albatrosses are seabirds in the genus ''Diomedea'' in the albatross family. The genus ''Diomedea'' formerly included all albatrosses except the sooty albatrosses, but in 1996 the genus was split, with the mollymawks and the North Pac ...
es, ''Diomedea'', but a study of their
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DN ...
showed that they are a
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic grou ...
taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
related to the sooty albatrosses, and they were placed in their own genus.


Taxonomy

The genus ''Thalassarche'' was introduced in 1853 by the German naturalist
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist and ornithologist. It was he who first requested Leopold Blaschka to make a set of glass marine invertebrate models for scientific education and museum ...
with the black-browed albatross as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
. The genus name combines the Ancient Greek ''thalassa'' meaning "sea" and ''arkhē'' meaning "power" or "command" (from ''arkhō'', to govern). The word ''mollymawk'', which dates to the late 17th century, comes from the
Dutch Dutch commonly refers to: * Something of, from, or related to the Netherlands * Dutch people () * Dutch language () Dutch may also refer to: Places * Dutch, West Virginia, a community in the United States * Pennsylvania Dutch Country People ...
''mallemok'', which means ''mal'' – foolish and ''mok'' – gull. Another etymology suggests that it comes from the German Mallemugge, a term used originally for midges or flies that whirled about lights. Mollymawks are a type of albatross that belong to the family
Diomedeidae Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Paci ...
of the order
Procellariiformes Procellariiformes is an order of seabirds that comprises four families: the albatrosses, the petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still called tubenoses in English, procellariiforms are of ...
, along with
shearwater Shearwaters are medium-sized long-winged seabirds in the petrel family Procellariidae. They have a global marine distribution, but are most common in temperate and cold waters, and are pelagic outside the breeding season. Description These tu ...
s,
fulmar The fulmars are tubenosed seabirds of the family Procellariidae. The family consists of two extant species and two extinct fossil species from the Miocene. Fulmars superficially resemble gulls, but are readily distinguished by their flight o ...
s, storm-petrels, and
diving-petrel The diving petrels form a genus, ''Pelecanoides'', of seabirds in the family Procellariidae. There are four very similar species of diving petrels, distinguished only by small differences in the coloration of their plumage, habitat, and bill co ...
s. They share certain identifying features. First, they have nasal passages that attach to the upper bill called
naricorns A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called tu ...
, although the nostrils are on the sides of the bill. The bills of Procellariiformes are also unique in that they are split into between seven and nine horny plates. They also have a
salt gland The salt gland is an organ for excreting excess salts. It is found in the cartilaginous fishes subclass elasmobranchii (sharks, rays, and skates), seabirds, and some reptiles. Salt glands can be found in the rectum of sharks. Birds and rept ...
that is situated above the nasal passage and helps desalinate their bodies, due to the high amount of ocean water that they imbibe. It excretes a high saline solution from their nose. Finally, they produce a
stomach oil Stomach oil is the light oil composed of neutral dietary lipids found in the proventriculus (fore-gut) of birds in the order Procellariiformes. All albatrosses, procellarids (gadfly petrels and shearwaters) and northern and austral storm petrel ...
made up of
wax ester A wax ester (WE) is an ester of a fatty acid and a fatty alcohol. Wax esters comprise the main components of three commercially important waxes: carnauba wax, candelilla wax, and beeswax.. Wax esters are formed by combining one fatty acid with o ...
s and
triglycerides A triglyceride (TG, triacylglycerol, TAG, or triacylglyceride) is an ester derived from glycerol and three fatty acids (from ''tri-'' and '' glyceride''). Triglycerides are the main constituents of body fat in humans and other vertebrates, a ...
that is stored in the
proventriculus The proventriculus is part of the digestive system of birds.Encarta World English Dictionary orth American Edition(2007). ''Proventriculus''. Source: (accessed: December 18, 2007) An analogous organ exists in invertebrates and insects. Birds The ...
. This can be sprayed out of their mouths as a defence against predators as well as an energy rich food source for chicks and for the adults during their long flights. The
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
species ''Thalassarche thyridata'' known from a skull fragment from the Late
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent ...
of
Victoria, Australia Victoria is a state in southeastern Australia. It is the second-smallest state with a land area of , the second most populated state (after New South Wales) with a population of over 6.5 million, and the most densely populated state in A ...
shows that the genus had already diverged from the sooty albatrosses 10 mya.


Species


Extant Species

The genus contains nine species:


Fossils

* '' Thalassarche thyridata''


Description

Mollymawks have the largest range in size of all the albatross genera, as their wingspans are . Mollymawks have what has been described as
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st centur ...
-like
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
, with dark black backs, mantle and tails and lighter heads, underwings and bellies. The heads of several species are often slightly darker grey, or have dark around the eyes. They all have a colourful pinkish flesh stripe from their
gape The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food, ...
to their ear that is shown during displays. They have distinctive bill structure and colouring which makes for easier identifying than other albatrosses. The bills of mollymawks are either brightly coloured orange or yellow, or dark with several bright yellow lines.


See also

*
List of albatross breeding locations The following is a list of locations where albatrosses breed, together with a list of the species found at each location. North Pacific Ocean *Bonin Islands - black-footed, Laysan *French Frigate Shoals - Laysan, black-footed *Guadalupe Island ...


References


Further reading

*


External links

* * {{Authority control Thalassarche Taxa named by Ludwig Reichenbach