Two-lined Monocle Bream
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''Scolopsis bilineata'', the two-lined monocle bream, bridled monocle bream, bridled spinecheek, double-lined coral bream or yellow-finned spine-cheek, is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
belonging to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Nemipteridae, the threadfin breams. This species is found in the Indian and Western Pacific Oceans.


Taxonomy

''Scolopsis bilineata'' was first formally described as ''Anthias bilineatus'' by the German
physician A physician, medical practitioner (British English), medical doctor, or simply doctor is a health professional who practices medicine, which is concerned with promoting, maintaining or restoring health through the Medical education, study, Med ...
and
naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
Marcus Elieser Bloch Marcus Elieser Bloch (1723–1799) was a German physician and naturalist who is best known for his contribution to ichthyology through his multi-volume catalog of plates illustrating the fishes of the world. Brought up in a Hebrew-speaking Jewish ...
in 1793 with its type locality given as Japan. The 5th edition of ''
Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...
'' classifies the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Scolopsis'' within the family Nemipteridae which it places in the
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Spariformes Spariformes is an order of ray-finned fishes consisting of six families within the series Percomorpha. Taxonomy Spariformes was first used as a taxonomic term in 1860 by the Dutch physician, herpetologist and ichthyologist Pieter Bleeker. Trad ...
.


Etymology

''Scolopsis bilineata'' has the
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''bilineata'' which means "two lined", a reference to its the two curved, parallel dark stripes on the head of adults.


Description

''Scolopsis bilineata'' has its
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
supported by 10 spines and 7 soft rays while the
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
contains 3 spines and 7 soft rays. Its body has a depth that fits into its
standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...
between 2.5 and 3 times with the length of the snout being less than the diameter of the eye. The scales on the head reach forward as far as or beyond the front nostrils and there are scales on the lower limb of the
preoperculum This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes. A B C ...
. There is a forward pointing spine on the suborbital. The
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...
s are long, extending to or nearly to the level of the origin of the anal fin. The caudal fin is forked. There is white stripe with black margins that arcs upwards from underneath the eye to the centre of the base of the dorsal fin. There is a pair of white lines over and to the rear of the eyes. The spiny part of the dorsal fin is yellow and there is a white area on the back underneath the soft-rayed part of the dorsal fin. In the juveniles the upper half of body is black with two yellow stripes and the middle of the flanks have a wide poorly defined white stripe with a black mark on the anal fin. This species has a maximum published
total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...
of although
standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...
is more typical.


Distribution and habitat

''Scolopsis bilineata'' is found in the eastern Indian Ocean and the western Pacific Ocean. It ranges from the Maldives and Sri Lanka east as far as Fiji and the
Caroline Islands The Caroline Islands (or the Carolines) are a widely scattered archipelago of tiny islands in the western Pacific Ocean, to the north of New Guinea. Politically, they are divided between the Federated States of Micronesia (FSM) in the cen ...
, it has also been reported from Tonga. Its range extends north to the
Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Geography of Taiwan, Taiwan: the Ryukyu Islands are divided into the Satsunan Islands (Ōsumi Islands, Ōsumi, Tokara Islands, Tokara and A ...
and south to Australia. Its Australian range extends from the
Houtman Abrolhos The Houtman Abrolhos (often called the Abrolhos Islands) is a chain of 122 islands and associated coral reefs in the Indian Ocean off the west coast of Australia about west of Geraldton, Western Australia. It is the southernmost true coral r ...
in ]
Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...
along the western, northern and eastern coasts as far as
Sydney Sydney is the capital city of the States and territories of Australia, state of New South Wales and the List of cities in Australia by population, most populous city in Australia. Located on Australia's east coast, the metropolis surrounds Syd ...
in
New South Wales New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...
. It has also been recorded from the
Rowley Shoals The Rowley Shoals is a group of three atoll-like coral reefs south of the Timor Sea, about west of Broome on the northwestern Australian coast, centred on , on the edge of one of the widest continental shelves in the world. Each atoll covers a ...
and Scott Reef off Western Australia, the Ashmore Reef in the
Timor Sea The Timor Sea (, , or ) is a relatively shallow sea in the Indian Ocean bounded to the north by the island of Timor with Timor-Leste to the north, Indonesia to the northwest, Arafura Sea to the east, and to the south by Australia. The Sunda Tr ...
,
Christmas Island Christmas Island, officially the Territory of Christmas Island, is an States and territories of Australia#External territories, Australian external territory in the Indian Ocean comprising the island of the same name. It is about south o ...
,
Cocos (Keeling) Islands The Cocos (Keeling) Islands (), officially the Territory of Cocos (Keeling) Islands (; ), are an Australian external territory in the Indian Ocean, comprising a small archipelago approximately midway between Australia and Sri Lanka and rel ...
, Elizabeth and Middleton Reefs Marine National Park Reserve and
Lord Howe Island Lord Howe Island (; formerly Lord Howe's Island) is an irregularly crescent-shaped volcanic remnant in the Tasman Sea between Australia and New Zealand, part of the Australian state of New South Wales. It lies directly east of mainland Port ...
. It is found at depths between on coral reefs, with the juveniles being found inshore, in lagoons or in areas of rubble.


Biology

''Scolopsis bilineata'' are found typically either as solitary individuals or in pairs, frequently gather in small groups. Its prey is mainly benthic invertebrates and smaller fishes. It is a protogynous hermaphrodite but there is no evidence that this is functional and the sex change occur prior to sexual maturation. The juveniles are Batesian mimics of the linespot fangblenny ('' Meiacanthus grammistes''), a species armed with venomous canin-like teeth. The two-lined monocle bream exhibits
biofluorescence Biofluorescence is fluorescence exhibited by a living organism: part of the organism absorbs light or other radiation at one wavelength and emits visible light at another, usually longer. The absorbed radiation is often blue or ultraviolet, while t ...
, that is, when illuminated by blue or ultraviolet light, it re-emits it as green, and appears differently than under white light illumination (only stripes on the upper front part are visible). Biofluorescence may assist in intraspecific communication and camouflage, blending the fish with green-fluorescing ''
Acropora ''Acropora'' is a genus of small polyp stony coral in the phylum Cnidaria. Some of its species are known as table coral, elkhorn coral, and staghorn coral. Over 149 species are described. ''Acropora'' species are some of the major reef corals ...
'' corals.


Fisheries

''Scolpsis bilineata'' is occasionally found in small numbers in fish markets and there is no fishery specifically targeting this species. In the Philippines this species is caught to be sold in the
aquarium An aquarium (: aquariums or aquaria) is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. fishkeeping, Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aquati ...
trade.


References


External links

* * * Two-line Monocle Bream @ Fishes of Australia
/small> {{Taxonbar, from=Q2415683 bilineata Fish of the Indian Ocean Fish described in 1793