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In
physical geography Physical geography (also known as physiography) is one of the three main branches of geography. Physical geography is the branch of natural science which deals with the processes and patterns in the natural environment such as the atmosphere, h ...
, a tundra () is a type of
biome A biome () is a distinct geographical region with specific climate, vegetation, and animal life. It consists of a biological community that has formed in response to its physical environment and regional climate. In 1935, Tansley added the ...
where tree growth is hindered by frigid temperatures and short growing seasons. There are three regions and associated types of tundra:
Arctic The Arctic (; . ) is the polar regions of Earth, polar region of Earth that surrounds the North Pole, lying within the Arctic Circle. The Arctic region, from the IERS Reference Meridian travelling east, consists of parts of northern Norway ( ...
, Alpine, and
Antarctic The Antarctic (, ; commonly ) is the polar regions of Earth, polar region of Earth that surrounds the South Pole, lying within the Antarctic Circle. It is antipodes, diametrically opposite of the Arctic region around the North Pole. The Antar ...
. Tundra vegetation is composed of dwarf
shrubs A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
, sedges, grasses,
moss Mosses are small, non-vascular plant, non-vascular flowerless plants in the taxonomic phylum, division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Wilhelm Philippe Schimper, Schimp. 1879) may also refer to the parent group bryo ...
es, and
lichen A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
s. Scattered trees grow in some tundra regions. The ecotone (or ecological boundary region) between the tundra and the forest is known as the
tree line The tree line is the edge of a habitat at which trees are capable of growing and beyond which they are not. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually low ...
or timberline. The tundra soil is rich in
nitrogen Nitrogen is a chemical element; it has Symbol (chemistry), symbol N and atomic number 7. Nitrogen is a Nonmetal (chemistry), nonmetal and the lightest member of pnictogen, group 15 of the periodic table, often called the Pnictogen, pnictogens. ...
and
phosphorus Phosphorus is a chemical element; it has Chemical symbol, symbol P and atomic number 15. All elemental forms of phosphorus are highly Reactivity (chemistry), reactive and are therefore never found in nature. They can nevertheless be prepared ar ...
. The soil also contains large amounts of biomass and decomposed biomass that has been stored as
methane Methane ( , ) is a chemical compound with the chemical formula (one carbon atom bonded to four hydrogen atoms). It is a group-14 hydride, the simplest alkane, and the main constituent of natural gas. The abundance of methane on Earth makes ...
and
carbon dioxide Carbon dioxide is a chemical compound with the chemical formula . It is made up of molecules that each have one carbon atom covalent bond, covalently double bonded to two oxygen atoms. It is found in a gas state at room temperature and at norma ...
in the
permafrost Permafrost () is soil or underwater sediment which continuously remains below for two years or more; the oldest permafrost has been continuously frozen for around 700,000 years. Whilst the shallowest permafrost has a vertical extent of below ...
, making the tundra soil a
carbon sink A carbon sink is a natural or artificial carbon sequestration process that "removes a  greenhouse gas, an aerosol or a precursor of a greenhouse gas from the atmosphere". These sinks form an important part of the natural carbon cycle. An overar ...
. As global warming heats the ecosystem and causes soil thawing, the permafrost carbon cycle accelerates and releases much of these soil-contained greenhouse gases into the atmosphere, creating a feedback cycle that contributes to
global warming Present-day climate change includes both global warming—the ongoing increase in global average temperature—and its wider effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes ...
.


Etymology

The term is a Russian word adapted from the
Sámi languages The Sámi languages ( ), also rendered in English language, English as Sami and Saami, are a group of Uralic languages spoken by the Indigenous Sámi peoples in Northern Europe (in parts of northern Finland, Norway, Sweden, and extreme northwest ...
, specifically from the Kildin Sámi word ''tūndâr'', meaning "treeless plain".


Arctic

Arctic tundra occurs in the far
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
, north of the
taiga Taiga or tayga ( ; , ), also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches. The taiga, or boreal forest, is the world's largest land biome. In North A ...
belt. The word "tundra" usually refers only to the areas where the subsoil is
permafrost Permafrost () is soil or underwater sediment which continuously remains below for two years or more; the oldest permafrost has been continuously frozen for around 700,000 years. Whilst the shallowest permafrost has a vertical extent of below ...
, or permanently frozen soil. (It may also refer to the treeless plain in general so that northern Sápmi would be included.) Permafrost tundra includes vast areas of northern Russia and Canada. The polar tundra is home to several peoples who are mostly nomadic reindeer herders, such as the Nganasan and Nenets in the permafrost area (and the
Sami Acronyms * SAMI, ''Synchronized Accessible Media Interchange'', a closed-captioning format developed by Microsoft * Saudi Arabian Military Industries, a government-owned defence company * South African Malaria Initiative, a virtual expertise ne ...
in Sápmi). Arctic tundra contains areas of stark landscape and is frozen for much of the year. The soil there is frozen from down, making it impossible for trees to grow there. Instead, bare and sometimes rocky land can only support certain kinds of Arctic vegetation, low-growing plants such as moss, heath (
Ericaceae The Ericaceae () are a Family (biology), family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with about 4,250 known species spread acros ...
varieties such as crowberry and black bearberry), and
lichen A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
. There are two main seasons, winter and summer, in the polar tundra areas. During the winter it is very cold, dark, and windy with the average temperature around , sometimes dipping as low as . However, extreme cold temperatures on the tundra generally do not drop as low as those experienced in taiga areas further south (for example, Russia's, Canada's, and Alaska's lowest temperatures were recorded in locations south of the tree line). During the summer, temperatures rise somewhat, and the top layer of seasonally-frozen soil melts, leaving the ground very soggy. The tundra is covered in marshes, lakes, bogs, and streams during the warm months. Generally daytime temperatures during the summer rise to about but can often drop to or even below freezing. Arctic tundras are sometimes the subject of habitat conservation programs. In Canada and Russia, many of these areas are protected through a national Biodiversity Action Plan. Tundra tends to be windy, with winds often blowing upwards of . However, it is desert-like, with only about of precipitation falling per year (the summer is typically the season of maximum precipitation). Although precipitation is light, evaporation is also relatively minimal. During the summer, the permafrost thaws just enough to let plants grow and reproduce, but because the ground below this is frozen, the water cannot sink any lower, so the water forms the lakes and marshes found during the summer months. There is a natural pattern of accumulation of fuel and wildfire which varies depending on the nature of vegetation and terrain. Research in Alaska has shown fire-event return intervals (FRIs) that typically vary from 150 to 200 years, with dryer lowland areas burning more frequently than wetter highland areas. The
biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
of tundras is low: 1,700 species of vascular plants and only 48 species of land mammals can be found, although millions of birds migrate there each year for the marshes. There are also a few fish species. There are few species with large populations. Notable plants in the Arctic tundra include blueberry ('' Vaccinium uliginosum''), crowberry ('' Empetrum nigrum''), reindeer lichen (''
Cladonia rangiferina ''Cladonia rangiferina'', also known as reindeer cup lichen, reindeer lichen (cf. Swedish language, Sw. ''renlav'') or grey reindeer lichen, is a light-coloured fruticose lichen, fruticose, Cladonia, cup lichen species in the family Cladoniaceae. ...
''), lingonberry ('' Vaccinium vitis-idaea''), and Labrador tea ('' Rhododendron groenlandicum''). Notable animals include
reindeer The reindeer or caribou (''Rangifer tarandus'') is a species of deer with circumpolar distribution, native to Arctic, subarctic, tundra, taiga, boreal, and mountainous regions of Northern Europe, Siberia, and North America. It is the only re ...
(caribou), musk ox, Arctic hare,
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small species of fox native to the Arctic regions of the Northern Hemisphere and common throughout the Tundra#Arctic tundra, Arctic tundra biome. I ...
, snowy owl, ptarmigan, northern red-backed voles,
lemming A lemming is a small rodent, usually found in or near the Arctic in tundra biomes. Lemmings form the subfamily Arvicolinae (also known as Microtinae) together with voles and muskrats, which form part of the superfamily Muroidea, which also inclu ...
s, the
mosquito Mosquitoes, the Culicidae, are a Family (biology), family of small Diptera, flies consisting of 3,600 species. The word ''mosquito'' (formed by ''Musca (fly), mosca'' and diminutive ''-ito'') is Spanish and Portuguese for ''little fly''. Mos ...
, and even
polar bear The polar bear (''Ursus maritimus'') is a large bear native to the Arctic and nearby areas. It is closely related to the brown bear, and the two species can Hybrid (biology), interbreed. The polar bear is the largest extant species of bear ...
s near the ocean. Tundra is largely devoid of poikilotherms such as frogs or lizards. Due to the harsh climate of Arctic tundra, regions of this kind have seen little human activity, even though they are sometimes rich in natural resources such as
petroleum Petroleum, also known as crude oil or simply oil, is a naturally occurring, yellowish-black liquid chemical mixture found in geological formations, consisting mainly of hydrocarbons. The term ''petroleum'' refers both to naturally occurring un ...
,
natural gas Natural gas (also fossil gas, methane gas, and gas) is a naturally occurring compound of gaseous hydrocarbons, primarily methane (95%), small amounts of higher alkanes, and traces of carbon dioxide and nitrogen, hydrogen sulfide and helium ...
, and
uranium Uranium is a chemical element; it has chemical symbol, symbol U and atomic number 92. It is a silvery-grey metal in the actinide series of the periodic table. A uranium atom has 92 protons and 92 electrons, of which 6 are valence electrons. Ura ...
. In recent times this has begun to change in
Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
, Russia, and some other parts of the world: for example, the Yamalo-Nenets Autonomous Okrug produces 90% of Russia's natural gas.


Relationship to climate change

A severe threat to tundra is
global warming Present-day climate change includes both global warming—the ongoing increase in global average temperature—and its wider effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes ...
, which causes
permafrost Permafrost () is soil or underwater sediment which continuously remains below for two years or more; the oldest permafrost has been continuously frozen for around 700,000 years. Whilst the shallowest permafrost has a vertical extent of below ...
to thaw. The thawing of the permafrost in a given area on human time scales (decades or centuries) could radically change which species can survive there. It also represents a significant risk to infrastructure built on top of permafrost, such as roads and pipelines. In locations where dead vegetation and
peat Peat is an accumulation of partially Decomposition, decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, Moorland, moors, or muskegs. ''Sphagnum'' moss, also called peat moss, is one of the most ...
have accumulated, there is a risk of wildfire, such as the of tundra which burned in 2007 on the north slope of the Brooks Range in Alaska. Such events may both result from and contribute to global warming. Carbon emissions from permafrost thaw contribute to the same warming which facilitates the thaw, making it a positive climate change feedback. The warming also intensifies Arctic
water cycle The water cycle (or hydrologic cycle or hydrological cycle) is a biogeochemical cycle that involves the continuous movement of water on, above and below the surface of the Earth across different reservoirs. The mass of water on Earth remains fai ...
, and the increased amounts of warmer rain are another factor which increases permafrost thaw depths. The
IPCC Sixth Assessment Report The Sixth Assessment Report (AR6) of the United Nations (UN) Intergovernmental Panel on Climate Change (IPCC) is the sixth in a series of reports which assess the available scientific information on climate change. Three Working Groups (WGI, II, ...
estimates that carbon dioxide and methane released from permafrost could amount to the equivalent of 14–175 billion tonnes of carbon dioxide per of warming. For comparison, by 2019, ''annual'' anthropogenic emission of carbon dioxide alone stood around 40 billion tonnes. A 2018 perspectives article discussing
tipping points in the climate system In Climatology, climate science, a tipping point is a critical threshold that, when crossed, leads to large, accelerating and often irreversible changes in the climate system. If tipping points are crossed, they are likely to have severe impac ...
activated around of global warming suggested that at this threshold, permafrost thaw would add a further to global temperatures by 2100, with a range of


Antarctic

Antarctic tundra occurs on Antarctica and on several Antarctic and subantarctic islands, including
South Georgia and the South Sandwich Islands South Georgia and the South Sandwich Islands (SGSSI) is a British Overseas Territory in the southern Atlantic Ocean. It is a remote and inhospitable collection of islands, consisting of South Georgia and a chain of smaller islands known as the ...
and the
Kerguelen Islands The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the subantarctic, sub-Antarctic region. They are among the Extremes on Earth#Remoteness, most i ...
. Most of Antarctica is too cold and dry to support vegetation, and most of the continent is covered by ice fields or cold deserts. However, some portions of the continent, particularly the
Antarctic Peninsula The Antarctic Peninsula, known as O'Higgins Land in Chile and Tierra de San Martin in Argentina, and originally as Graham Land in the United Kingdom and the Palmer Peninsula in the United States, is the northernmost part of mainland Antarctica. ...
, have areas of rocky soil that support plant life. The flora presently consists of around 300–400 species of lichens, 100 mosses, 25
liverworts Liverworts are a group of non-vascular plant, non-vascular embryophyte, land plants forming the division Marchantiophyta (). They may also be referred to as hepatics. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in wh ...
, and around 700 terrestrial and aquatic algae species, which live on the areas of exposed rock and soil around the shore of the continent. Antarctica's two flowering plant species, the Antarctic hair grass (''Deschampsia antarctica'') and
Antarctic pearlwort ''Colobanthus quitensis'', also known as the Antarctic pearlwort, is one of two native flowering plants found in the Antarctic region, the other being Deschampsia antarctica, Antarctic hair grass. It has yellow flowers and grows about tall, givi ...
(''Colobanthus quitensis''), are found on the northern and western parts of the Antarctic Peninsula. In contrast with the Arctic tundra, the Antarctic tundra lacks a large mammal fauna, mostly due to its physical isolation from the other continents. Sea mammals and sea birds, including seals and penguins, inhabit areas near the shore, and some small mammals, like rabbits and cats, have been introduced by humans to some of the subantarctic islands. The Antipodes Subantarctic Islands tundra
ecoregion An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...
includes the Bounty Islands, Auckland Islands,
Antipodes Islands The Antipodes Islands (, ) are inhospitable and uninhabited volcanic islands in subantarctic waters to the south of – and territorially part of – New Zealand. The archipelago lies to the southeast of Stewart Island / Rakiura, and to the ...
, the Campbell Island group, and
Macquarie Island Macquarie Island is a subantarctic island in the south-western Pacific Ocean, about halfway between New Zealand and Antarctica. It has been governed as a part of Tasmania, Australia, since 1880. It became a Protected areas of Tasmania, Tasmania ...
. Species endemic to this ecoregion include ''
Corybas dienemus ''Corybas dienemus'', commonly known as the windswept helmet-orchid, is one of two Corybas (plant), helmet orchids endemism, endemic to Australia's subantarctic Macquarie Island, and the first orchid to be found there. It is a relatively small ...
'' and '' Corybas sulcatus'', the only subantarctic orchids; the royal penguin; and the Antipodean albatross. There is some ambiguity on whether Magellanic moorland, on the west coast of Patagonia, should be considered tundra or not. Phytogeographer Edmundo Pisano called it tundra () since he considered the low temperatures key to restrict plant growth. More recent approaches have since recognized it as a temperate grassland, restricting southern tundra to coastal Antarctica and its islands. The flora and fauna of Antarctica and the Antarctic Islands (south of 60° south latitude) are protected by the Antarctic Treaty.


Alpine

Alpine tundra does not contain trees because the climate and soils at high altitude block tree growth. The cold climate of the alpine tundra is caused by the low air temperatures, and is similar to
polar climate The polar climate regions are characterized by a lack of warm summers but with varying winters. Every month a polar climate has an average temperature of less than . Regions with a polar climate cover more than 20% of the Earth's area. Most of ...
. Alpine tundra is generally better drained than arctic soils. Alpine tundra transitions to subalpine forests below the tree line; stunted forests occurring at the forest-tundra ecotone (the
treeline The tree line is the edge of a habitat at which trees are capable of growing and beyond which they are not. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually low ...
) are known as '' Krummholz''. Alpine tundra can be affected by woody plant encroachment. Alpine tundra occurs in mountains worldwide. The flora of the alpine tundra is characterized by plants that grow close to the ground, including perennial grasses, sedges, forbs, cushion plants,
moss Mosses are small, non-vascular plant, non-vascular flowerless plants in the taxonomic phylum, division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Wilhelm Philippe Schimper, Schimp. 1879) may also refer to the parent group bryo ...
es, and
lichen A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
s. The flora is adapted to the harsh conditions of the alpine environment, which include low temperatures, dryness, ultraviolet radiation, and a short growing season.


Climatic classification

Tundra climates ordinarily fit the
Köppen climate classification The Köppen climate classification divides Earth climates into five main climate groups, with each group being divided based on patterns of seasonal precipitation and temperature. The five main groups are ''A'' (tropical), ''B'' (arid), ''C'' (te ...
''ET'', signifying a local climate in which at least one month has an average temperature high enough to melt snow (), but no month with an average temperature in excess of . The cold limit generally meets the ''EF'' climates of permanent ice and snows; the warm-summer limit generally corresponds with the poleward or altitudinal limit of trees, where they grade into the
subarctic climate The subarctic climate (also called subpolar climate, or boreal climate) is a continental climate with long, cold (often very cold) winters, and short, warm to cool summers. It is found on large landmasses, often away from the moderating effects of ...
s designated ''Dfd'', ''Dwd'' and ''Dsd'' (extreme winters as in parts of
Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
), ''Dfc'' typical in Alaska, Canada, mountain areas of
Scandinavia Scandinavia is a subregion#Europe, subregion of northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. ''Scandinavia'' most commonly refers to Denmark, Norway, and Sweden. It can sometimes also ...
,
European Russia European Russia is the western and most populated part of the Russia, Russian Federation. It is geographically situated in Europe, as opposed to the country's sparsely populated and vastly larger eastern part, Siberia, which is situated in Asia ...
, and
Western Siberia Western Siberia or West Siberia ( rus, Западная Сибирь, p=ˈzapədnəjə sʲɪˈbʲirʲ; , ) is a region in North Asia. It is part of the wider region of Siberia that is mostly located in the Russia, Russian Federation, with a Sout ...
(cold winters with months of freezing). Despite the potential diversity of climates in the ''ET'' category involving precipitation, extreme temperatures, and relative wet and dry seasons, this category is rarely subdivided. Rainfall and snowfall are generally slight due to the low
vapor pressure Vapor pressure or equilibrium vapor pressure is the pressure exerted by a vapor in thermodynamic equilibrium with its condensed phases (solid or liquid) at a given temperature in a closed system. The equilibrium vapor pressure is an indicat ...
of water in the chilly atmosphere, but as a rule potential evapotranspiration is extremely low, allowing soggy terrain of swamps and bogs even in places that get precipitation typical of deserts of lower and middle latitudes. The amount of native tundra biomass depends more on the local temperature than the amount of precipitation.


See also

* Alas * Fellfield * List of tundra ecoregions from the WWF * Mammoth steppe * Park Tundra


References


Further reading

* * * *


External links


WWF Tundra Ecoregions


* ttps://web.archive.org/web/20051225213602/http://www.antarctica.ac.uk/About_Antarctica/Wildlife/Plants/ British Antarctica Surveybr>Antarctica: West of the Transantarctic Mountains
{{Authority control Terrestrial biomes Geography of the Arctic Environment of the Arctic Köppen climate types Nearctic realm Palearctic realm