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''Tritylodon'' (from the Greek for "three-cusped tooth") is an extinct genus of tritylodonts, one of the most advanced group of
cynodont Cynodontia () is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 Megaannum, mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extin ...
therapsids. They lived in the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
and possibly Late Triassic periods along with
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s. They also shared many characteristics with
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s, and were once considered mammals because of overall skeleton construction. That was changed due to them retaining the vestigial amniote jawbones and a different skull structure. Tritylodonts are now regarded as non-mammalian
synapsid Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant rept ...
s.


Characteristics

If a living ''Tritylodon'' were to be seen today, it would look a lot like a large rodent. They were about long but there is no certainty about the exact weight. Their method of chewing food, a grinding motion with the bottom teeth sliding against the top teeth, resembled that of rodents as well. The bottom teeth were much like a set of cusps and the top teeth were a set of matching grooves that matched perfectly allowing this motion. There were large incisors at the very front of their mouth separated by a gap from the rest of the teeth. The incisors would stick out and remain slightly visible when the mouth was closed. The legs were directly beneath the body like mammals, unlike the earlier therapsids with sprawling limbs. These animals were burrowers; the structure of the shoulder, front limbs, and large front incisors show this. They used their incisors to help dig and unearth buried plants. The way they ate and the shape of their teeth demonstrate that ''Tritylodons'' were probably primarily
herbivorous A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
(though some tritylodontids show evidence of more omnivorous diets, and modern analogues like
rodents Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
tend to be more omnivorous than their dentitions lead on). Any of the Tritylodonts including ''Tritylodon'' were
warm-blooded Warm-blooded is a term referring to animal species whose bodies maintain a temperature higher than that of their environment. In particular, homeothermic species (including birds and mammals) maintain a stable body temperature by regulating ...
or endothermic. Like most non-
placental Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguished ...
mammalimorphs, it had
epipubic Epipubic bones are a pair of bones projecting forward from the pelvic bones of modern marsupials, monotremes and fossil mammals like multituberculates, and even basal eutherians (the ancestors of placentals, who lack them). They first occur in ...
bones, aiding in its erect gait but preventing the expansion of the abdomen, making it unable to go through prolonged pregnancy and instead give birth to larval young like modern
marsupials Marsupials are a diverse group of mammals belonging to the infraclass Marsupialia. They are natively found in Australasia, Wallacea, and the Americas. One of marsupials' unique features is their reproductive strategy: the young are born in a ...
and
monotremes Monotremes () are mammals of the order Monotremata. They are the only group of living mammals that lay eggs, rather than bearing live young. The extant monotreme species are the platypus and the four species of echidnas. Monotremes are typified ...
.


Habitat

The ''Tritylodons habitat was limited to the forests of South Africa When the species originated, about 200 million years ago, the African area was drier and hotter. But for most of their existence the climate was tropical and wetter.


Fossils

The ''Tritylodon'' fossils in South Africa are found concentrated mainly in an area about 11,000 km2 (4,250 mi2). They have been found in floodplain deposits of the Lower Jurassic
Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, so ...
(upper
Karoo Supergroup The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a per ...
). In this area there have been so many findings it has been named the Tritylodon Acme Zone. The fossil findings have all been in the Free State of South Africa. The genus ''Tritylodon'' of the Tritylodonts is restricted to the South African forms: ''Tritylodon longaevus'' and ''Tritylodon maximus''. It is suggested that ''T. maximus'' is either a large ''T. longeavus'' or a closely related species. If it is a closely related species it could possibly be ecological succession since the larger ''T. maximus'' fossils have been dated in the Sinemurian–Pliensbachian mainly less than 190 million years ago and the ''T. longaevus'' in the Hettangian–Sinemurian mainly more than 190 million years ago. With the fossil findings of each species overlapping in Sinemurian stage, the fossils show two differences, ''T. maximus'' being larger and having nine upper postcanines (neither species had canine teeth) instead of the seven teeth like ''T. longeavus''. All other structures of the two ''Tritylodon'' species were the same.http://home.arcor.de/ktdykes/jtherap.htm#tritylodon Below is a
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships:


References


External links

* * * * *https://archive.today/20130204222846/http://tritylodontidae.totallyexplained.com/ * {{Taxonbar, from=Q1441437 Tritylodontidae Early Jurassic synapsids of Africa Fossil taxa described in 1884 Taxa named by Richard Owen