Tritheledontid
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Tritheledontidae, the tritheledontids or ictidosaurs, is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of small to medium-sized (about 10 to 20 cm long)
cynodont Cynodontia () is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 Megaannum, mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extin ...
s. They were highly
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
-like, specialized cynodonts, although they still retained a few
reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
-like anatomical traits. Tritheledontids were mainly
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
or
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores we ...
, though some species may have developed omnivory. Their skeletons show that they had a close relationship to mammals. Tritheledontids or their closest relatives may have given rise to the
mammaliaforms Mammaliaformes ("mammalian forms") is a clade of synapsid tetrapods that includes the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts during the Late Triassic. It is defined as ...
. The tritheledontids were one of the longest lived non-mammalian therapsid lineages, living from the late
Triassic The Triassic ( ; sometimes symbolized šŸˆ) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
to the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
period. Tritheledontids became extinct in the Jurassic period, possibly due to competition with
prehistoric mammal This is an incomplete list of prehistoric mammals. It does not include extant mammals or recently extinct mammals. For extinct primate species, see: list of fossil primates.Mikko's Phylogeny Archiv Mammaliaformes ' *Genus †''Adelobasile ...
s such as the
eutriconodont Eutriconodonta is an order (biology), order of early mammals. Eutriconodonts existed in Asia (including Insular India, pre-contact India), Africa, Europe, North America, North and South America during the Jurassic and the Cretaceous periods. The ...
s. They are known from finds in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
and
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
, indicating that they may have lived only on the
supercontinent In geology, a supercontinent is the assembly of most or all of Earth's continent, continental blocks or cratons to form a single large landmass. However, some geologists use a different definition, "a grouping of formerly dispersed continents", ...
of
Gondwana Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The remnants of Gondwana make up around two-thirds of today's continental area, including South America, Africa, Antarctica, Australia (continent), Australia, Zea ...
. The family Tritheledontidae was named by South African paleontologist
Robert Broom Robert Broom Fellow of the Royal Society, FRS FRSE (30 November 1866 6 April 1951) was a British- South African medical doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University ...
in 1912. The family is often misspelled "Trithelodontidae". It is possible that tritheledontids had vibrissae, according to the
PBS The Public Broadcasting Service (PBS) is an American public broadcaster and non-commercial, free-to-air television network based in Arlington, Virginia. PBS is a publicly funded nonprofit organization and the most prominent provider of educat ...
documentary, ''Your Inner Fish''. A common ancestor of all therian mammals did so. It is possible that the development of the whisker sensory system played an important role in mammalian development, more generally.


Phylogeny

Cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
after Ruta, Botha-Brink, Mitchell and Benton (2013):


See also

*
Evolution of mammals The evolution of mammals has passed through many stages since the first appearance of their synapsid ancestors in the Pennsylvanian (geology), Pennsylvanian sub-period of the late Carboniferous period. By the mid-Triassic, there were many synaps ...


References


External links


PBS YouTube animated video
of a tritheledontid {{Taxonbar, from1=Q21646750, from2=Q140837 Late Triassic first appearances Middle Jurassic extinctions Prehistoric therapsid families