Trachilos Footprints
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The Trachilos footprints are possibly
tetrapod A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...
footprints which show hominin-like characteristics from the late
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
on the western
Crete Crete ( ; , Modern Greek, Modern: , Ancient Greek, Ancient: ) is the largest and most populous of the Greek islands, the List of islands by area, 88th largest island in the world and the List of islands in the Mediterranean#By area, fifth la ...
, close to the village of Trachilos, west of
Kissamos Kissamos () is a town and a municipality in the west of the island of Crete, Greece. It is part of the Chania regional unit and of the former Kissamos Province which covers the northwest corner of the island. The town of Kissamos is also known as ...
, in the
Chania Prefecture Chania (), also spelled Hania, is one of the four regional units of Crete; it covers the westernmost quarter of the island. Its capital is the city of Chania. Chania borders only one other regional unit: that of Rethymno to the east. The western ...
. Researchers describe the tracks as representing at least one apparent bipedal
hominin The Hominini (hominins) form a taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos), and in standard usage exclude the genus '' Gorilla'' ( gorillas) ...
or an unknown
primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...
. The stratum in which the footprints were found was dated to about 5.7 million years ago, which predates the previously earliest discovered hominin footprints by about two million years. Later studies show that the footprints might be more than 6 million years old. The researchers of the tracks suggest that it may imply the possibility of hominin evolution outside of Africa, contrary to the current
theory A theory is a systematic and rational form of abstract thinking about a phenomenon, or the conclusions derived from such thinking. It involves contemplative and logical reasoning, often supported by processes such as observation, experimentation, ...
. However, there is no consensus that these impressions are distinct enough to confidently assign to a primate or even a
vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...
, or that they are indeed footprints at all.


Discovery

The tracks were originally discovered by Gerard D. Gierliński, from the Polish Research Institute in
Warsaw Warsaw, officially the Capital City of Warsaw, is the capital and List of cities and towns in Poland, largest city of Poland. The metropolis stands on the Vistula, River Vistula in east-central Poland. Its population is officially estimated at ...
in 2002. During a visit to Trachilos on Crete, Gierliński found the tracks, and as he was not planning on staying in Trachilos, he recorded the footprints to investigate them in the future. In 2012 Gierliński received permission from the Greek government to research the area, returning to Trachilos with other researchers to explore the tracks in detail. The researchers used methods such as laser scans and 3D imaging of the footprints, and compared them to apes and bears as well as humans.


Dates

The tracks were dated by using the underlying rock bed, predominantly
sedimentary Sedimentary rocks are types of rock formed by the cementation of sediments—i.e. particles made of minerals (geological detritus) or organic matter (biological detritus)—that have been accumulated or deposited at Earth's surface. Sedime ...
, and
foraminifera Foraminifera ( ; Latin for "hole bearers"; informally called "forams") are unicellular organism, single-celled organisms, members of a phylum or class (biology), class of Rhizarian protists characterized by streaming granular Ectoplasm (cell bio ...
. The study explains, "The coastal rocks at Trachilos lie within the Platanos Basin, and present a succession of shallow marine late Miocene carbonates and siliciclastics ..At the top, this marine succession terminates abruptly in the coarse-grained terrigenous sedimentary rocks of the Hellenikon Group The study continues, stating that the sedimentary rocks would have been created around 5.6 million years ago, at the time of the
Messinian salinity crisis In the Messinian salinity crisis (also referred to as the Messinian event, and in its latest stage as the Lago Mare event) the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (drying-up) throughout the latter part of ...
(mya). The researchers also found evidence of foraminifera, which were dated at 8.5 mya and 3.5 mya. Given the date of the sedimentary rocks and the foraminifera samples, the researches created an approximate interval of 8.5 mya to 5.6 mya. As the rock sediment containing the tracks resembled that of Hellenikon minerals, the tracks were estimated to be 5.7 mya within the given interval. In 2021, further research was published in regard to dating by Kirscher et al. Cyclostratigraphic data based on
magnetic susceptibility In electromagnetism, the magnetic susceptibility (; denoted , chi) is a measure of how much a material will become magnetized in an applied magnetic field. It is the ratio of magnetization (magnetic moment per unit volume) to the applied magnet ...
indicated that the Trachilos footprints are about 6.05 Ma old, which is 350,000 years older than was found previously. This puts the Trachilos footprints in the same period as the fossils of ''
Orrorin ''Orrorin'' is an extinct genus of primate within Homininae from the Miocene Lukeino Formation and Pliocene Mabaget Formation, both of Kenya. The type species is ''O. tugenenesis'', named in 2001, and a second species, ''O. praegens'', assigne ...
tugenensis'' from
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
.


Characteristics

The footprints were measured to range at 94-223 mm (3.7-8.8 inches) long and determined to be oriented in a south-west direction. There are clear pressure indexes, resembling that of a modern ''Homo sapiens''
plantigrade 151px, Portion of a human skeleton, showing plantigrade habit In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. ...
structure. The researchers also determined the presence of five digits in the imprints, classifying the track maker as pentadactyl, and lacking claws. As there was no visible evidence of forelimbs from the tracks, the track maker was identified as bipedal. Through 3D printing and laser scanning, there are impressions found which indicate a ball region, a pulling up motion of the foot, a
hallux Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being ''digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being ''plantigrade''; ...
, and small possible gaps between the first and other digit impressions. Poorly preserved prints lack these gaps, however. The lateral digit impressions become progressively smaller so that the digital region as a whole is strongly asymmetrical. The impression of the
hallux Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being ''digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being ''plantigrade''; ...
has a narrow neck and bulbous asymmetrical distal pad, indicating that the tracks were entaxonic. Morphometric analysis showed the footprints to have outlines that are distinct from modern non-hominin primates and resemble those of hominins. While younger than fossil records of hominins such as ''
Sahelanthropus ''Sahelanthropus'' is an extinct genus of hominid dated to about during the Late Miocene. The type species, ''Sahelanthropus tchadensis'', was first announced in 2002, based mainly on a partial cranium, nicknamed ''Toumaï'', discovered in north ...
'', found in
Chad Chad, officially the Republic of Chad, is a landlocked country at the crossroads of North Africa, North and Central Africa. It is bordered by Libya to Chad–Libya border, the north, Sudan to Chad–Sudan border, the east, the Central Afric ...
and dated around seven million years ago, the discovery potentially challenges the generally accepted theory that all early hominins were only present in Africa. The print morphology suggests that the trackmaker could be a basal member of the clade Hominini, but as Crete is some distance outside the known geographical range of pre-
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
hominins, researchers say that there is also a possibility that they represent a hitherto unknown late
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
primate that convergently evolved human-like foot anatomy.


News and controversy

When Gierliński and his team tried to publish the study, they received harsh criticism due to the findings going against the widely-accepted theory of early hominins evolving in
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
alone. According to the study, the Trachilos footprints may represent an early hominin or primate species that may have evolved hominin-like feet independently, outside of Africa. It also suggests the possibility of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
, wherein unrelated species adapt similar traits and characteristics to each other. Although convergent evolution is not an implausible explanation of the hominin-like trackways,, strong evidence is needed to support this interpretation. There were doubts if the tracks were footprints at all. This resulted in rejections from many scientific journals when offered to publish the study's findings. In an interview at the
CBC News CBC News is the division of the Canadian Broadcasting Corporation responsible for the news gathering and production of news programs on the corporation's English-language operations, namely CBC Television, CBC Radio, CBC News Network, and CBC ...
, researchers claimed that while they were trying to publish their work about the footprints at high-profile publications they got "ferociously aggressive responses", criticism and rejection from reviewers and editors. According to the researchers, “Basically, it wasn't a true
peer review Peer review is the evaluation of work by one or more people with similar competencies as the producers of the work (:wiktionary:peer#Etymology 2, peers). It functions as a form of self-regulation by qualified members of a profession within the ...
process at all,” “They were just trying to shut us down.” After multiple rejections from other publications, the study was eventually published in the journal, "Proceedings of the Geologists' Association." Shortly after the research about the footprints was published, eight prints were chiseled out of the rock and stolen. According to the newspaper '' Proto Thema'', the culprit was a high school teacher, who was later arrested by Crete authorities at Kasteli, Chania.Chung, Emily (February 23, 2018)
"One Hell of an Impression"
''CBC News''.
The prints were later found in his house and on a farm. David Begun, a paleoanthropologist at the
University of Toronto The University of Toronto (UToronto or U of T) is a public university, public research university whose main campus is located on the grounds that surround Queen's Park (Toronto), Queen's Park in Toronto, Ontario, Canada. It was founded by ...
, Canada, argues that the footprints might not necessarily belong to a human ancestor, despite its appearance. Similarly, Robin Crompton, a biological anthropologist at the
University of Liverpool The University of Liverpool (abbreviated UOL) is a Public university, public research university in Liverpool, England. Founded in 1881 as University College Liverpool, Victoria University (United Kingdom), Victoria University, it received Ro ...
, England, believes that although the footprints belong to a bipedal organism, they might not be human ancestors but “made by a member of the great ape clade”. Multiple fossilized footprints must be found and analyzed in order to accurately categorize the tracks by species. Meldrum and Sarmiento (2018) who studied the Trachilos prints ( ichnites) did not agree with Gierlinski et al. (2017) that these ichnites were made by a primate or even a vertebrate. Principally, Gierlinski et al. did not provide objective criteria for identifying prints as primate prints. The prints show no consistent and repetitive details that could identify them as being made by a primate, or even an animal with bilateral symmetry. Fatally, Gierlinski et al. did not explore alternative agents that could have produced the prints and provided no explanation as to how their analyses of print outlines deals with missing data and comparisons of non-homologous print outlines. Meldrum and Sarmiento concluded that there was insufficient evidence to argue the prints were made by an animal with bilateral symmetry. The Trachilos ichnites could have been made by non-vertebrate lifeforms or could even be the result of non-organic agents. Willem Jan Zachariasse and Lucas J. Lourens reestimated the footprints as to have originated at the Late Pliocene (3 mya) and question if hominids had created the footprints or if they were footprints. They argue that the shallow marine setting could not have been travelled across by hominins or bipedal primates but "in case they did, wave action must have erased them unless the tracks were made at the high tide line during a low sea level stand at the climax of one of the glacial periods that occurred every 41 kyr in this time span". Further, they state "Crete was an island since the Late
Tortonian The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ago). It follows the Serravallian and is followed by the Messinian. The Tort ...
but the present outline and geographic setting dates from the Late Pliocene with the deep Levantine Basin to the south and the marine South Aegean Basin (SAB) to the north. The latter basin formed in the Late Miocene by slab roll-back driven extension of the Aegean
lithosphere A lithosphere () is the rigid, outermost rocky shell of a terrestrial planet or natural satellite. On Earth, it is composed of the crust and the lithospheric mantle, the topmost portion of the upper mantle that behaves elastically on time ...
and was ~ 1000 m deep north of Crete (DSDP Site 378) during the Late Pliocene. The Late Pliocene SAB separated Crete from mainland Greece and Turkey by stretches of deep water of minimally 100 km wide according to the time–space reconstructions of the Aegean lithosphere in Van Hinsbergen and Schmid. Occurrences of deep marine Late Pliocene sediments on the islands of Kythira in between Crete and the Peloponnese, and Karpathos in between Crete and Turkey prove that these islands could not have been used as steppingstones for biped dispersal, let alone that they have been part of land bridges".


See also

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Convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
*
Early expansions of hominins out of Africa Several expansions of populations of archaic humans (genus ''Homo'') out of Africa and throughout Eurasia took place in the course of the Lower Paleolithic, and into the beginning Middle Paleolithic, between about 2.1 million and 0.2 million ye ...
*
Messinian salinity crisis In the Messinian salinity crisis (also referred to as the Messinian event, and in its latest stage as the Lago Mare event) the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (drying-up) throughout the latter part of ...


References

{{reflist Fossil trackways Archaeological discoveries in Crete Prehistoric Greece Crete Human evolution Hominin fossils