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A tracheid is a long and tapered lignified cell in the
in cross section
an
both in pine. {{Authority control Plant anatomy Plant physiology
A tracheid is a long and tapered lignified cell in the xylem
Xylem is one of the two types of transport tissue (biology), tissue in vascular plants, the other being phloem; both of these are part of the vascular bundle. The basic function of the xylem is to transport water upward from the roots to parts o ...
of vascular plants. It is a type of conductive cell called a tracheary element. Angiosperms also use another type of conductive cell, called vessel element
A vessel element or vessel member (also called a xylem vessel) is one of the cell types found in xylem, the water conducting tissue of plants. Vessel elements are found in most angiosperms (flowering plants) and in some gymnosperms such as cyca ...
s, to transport water through the xylem. The main functions of tracheid cells are to transport water and inorganic salts, and to provide structural support for trees. There are often pits on the cell walls of tracheids, which allows for water flow between cells. Tracheids are dead at functional maturity and do not have a protoplast
Protoplast (), is a biology, biological term coined by Johannes von Hanstein, Hanstein in 1880 to refer to the entire cell, excluding the cell wall. Protoplasts can be generated by stripping the cell wall from plant, bacterium, bacterial, or f ...
. The wood
Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
(softwood
Scots pine, a typical and well-known softwood
Softwood is wood from gymnosperm trees such as conifers. The term is opposed to hardwood, which is the wood from angiosperm trees. The main differences between hardwoods and softwoods is that the sof ...
) of gymnosperm
The gymnosperms ( ; ) are a group of woody, perennial Seed plant, seed-producing plants, typically lacking the protective outer covering which surrounds the seeds in flowering plants, that include Pinophyta, conifers, cycads, Ginkgo, and gnetoph ...
s such as pines and other conifer
Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
s is mainly composed of tracheids. Tracheids are also the main conductive cells in the primary xylem of fern
The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissue ...
s.
The tracheid was first named by the German botanist Carl Gustav Sanio in 1863, from the German ''Tracheide''.
Evolution
Tracheids were the main conductive cells found in early vascular plants. In the first 140–150 million years of vascular plant evolution, tracheids were the only type of conductive cells found in fossils of plant xylem tissues. Ancestral tracheids did not contribute significantly to structural support, as can be seen in extant ferns. Thefossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
record shows three different types of tracheid cells found in early plants, which were classified as S-type, G-type and P-type. The first two of them were lignified and had pores to facilitate the transportation of water between cells. The P-type tracheid cells had pits similar to extant plant tracheids. Later, more complex pits appeared, such as bordered pits on many tracheids, which allowed plants to transport water between cells while reducing the risk of cavitation and embolisms in the xylem.
As tracheids evolved along with secondary xylem tissues, specialized inter-tracheid pits appeared. Tracheid length and diameter also increased, with tracheid diameter increasing to an average length of 80 μm by the end of the Devonian
The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ...
period.
Tracheids then evolved into the vessel elements and structural fibers that make up angiosperm wood.
References
Further reading
*External links
*Pictures of softwood tracheidin cross section
an
both in pine. {{Authority control Plant anatomy Plant physiology