Toona Sinensis
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''Toona sinensis'', commonly called Chinese mahogany, Chinese cedar, Chinese toon, beef and onion plant, or red toon (; ; ; ; ) is a species of ''
Toona ''Toona'', commonly known as red cedar, toon (also spelled tun) or toona, tooni (in Nepal and India) is a genus in the mahogany family, Meliaceae, native from Afghanistan south to India, and east to North Korea, Papua New Guinea and eastern Austr ...
'' native to eastern and southeastern
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, ranging from northern Korean peninsula through most of eastern, central, and southwestern
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, in
Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
, northeastern
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
, Burma (
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
),
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
, and even present in
Malaysia Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
and western
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
.University of Melbourne
Sorting ''Toona'' names
/ref>Hong Kong trees

(in Chinese, with photos

Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .


Characteristics

It is a
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
growing to tall with a trunk up to 70 cm diameter. The
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
is brown, smooth on young trees, becoming scaly to shaggy on old trees. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...
, 50–70 cm long and 30–40 cm broad, with 10–40 leaflets, the terminal leaflet usually absent (paripinnate) but sometimes present (imparipennate); the individual leaflets 9–15 cm long and 2.5–4 cm broad, with an entire or weakly serrated margin. The young leaves are reddish-brown or purple, and have a smell. The
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s are produced in summer in
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
s 30–50 cm long at the end of a branch; each flower is small, 4–5 mm diameter, with five white or pale pink petals. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
is a golden capsule 2–3.5 cm long, containing several winged
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s.Taiwan Forestry
''Toona sinensis''
(in Chinese, with photos

It is similar to ''
Ailanthus altissima ''Ailanthus altissima'' ( ), commonly known as tree of heaven or ailanthus tree, is a deciduous tree in the quassia family. It is native to northeast and central China, and Taiwan. Unlike other members of the genus ''Ailanthus'', it is found ...
'' in appearance, but their leaves smell differently. Toona has rough bark, while ''A. altissima'' has smooth bark.


Cultivation

''Toona sinensis'' can reproduce both sexually and asexually, including seed propagation,
cutting Cutting is the separation or opening of a physical object, into two or more portions, through the application of an acutely directed force. Implements commonly used for wikt:cut, cutting are the knife and saw, or in medicine and science the sca ...
propagation and
tissue culture Tissue culture is the growth of tissue (biology), tissues or cell (biology), cells in an artificial medium separate from the parent organism. This technique is also called micropropagation. This is typically facilitated via use of a liquid, semi-s ...
. Seed propagation can provide large quantity of
seedling A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embry ...
s, which is suitable for the need of mass cultivation. Seeds soaked in warm water for a moderate amount of time before
sowing Sowing is the process of planting seeds. An area that has had seeds planted in it will be described as a sowed or sown area. Plants which are usually sown Among the major field crops, oats, wheat, and rye are sown, grasses and legumes are ...
are more likely to
germinate Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an flowering plant, angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the sp ...
. Normally, seeds of ''T. sinensis'' are sowed between the late March and early April in East Asia, and the time may vary depending on the actual planting area.
Sapling In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only pla ...
s grown from seeds in spring can be transplanted with leaves in fall.
Cutting Cutting is the separation or opening of a physical object, into two or more portions, through the application of an acutely directed force. Implements commonly used for wikt:cut, cutting are the knife and saw, or in medicine and science the sca ...
propagation uses a piece of stem or root of mature plant to grow a new plant in media like moist soil. This method has a higher survival rate in saplings than other methods. Usually semi-
lignified Lignin is a class of complex organic polymers that form key structural materials in the support tissues of most plants. Lignins are particularly important in the formation of cell walls, especially in wood and bark, because they lend rigidity an ...
stems are used in planting because those that have undergone full
lignification Lignin is a class of complex organic polymers that form key structural materials in the support tissues of most plants. Lignins are particularly important in the formation of cell walls, especially in wood and Bark (botany), bark, because they l ...
process are hard to take root or root slowly and stems that have not lignified are easy to decay after planting. NAA or
Vitamin D Vitamin D is a group of structurally related, fat-soluble compounds responsible for increasing intestinal absorption of calcium, magnesium, and phosphate, along with numerous other biological functions. In humans, the most important compo ...
solutions can help with the rooting of cut stems. Normally, ''T. sinensis'' stems are cut from mature plant and cultivated between late June and early July in East Asia, and time may differ depending on the actual planting area.
Tissue culture Tissue culture is the growth of tissue (biology), tissues or cell (biology), cells in an artificial medium separate from the parent organism. This technique is also called micropropagation. This is typically facilitated via use of a liquid, semi-s ...
of ''T. sinensis'' started from the late 1980s in China. Successful cases include the culture of ''T. sinensis'' seedlings on MS medium together with IAA and BA hormones. Since the 1980s, researchers have collected stems of mature ''T. sinensis'' trees from different regions and built a set of methods specifically for the tissue culture of rare varieties of ''Toona sinensis.''


Uses


Food and Nutrition

The young leaves of ''T. sinensis'' are extensively used as a
vegetable Vegetables are edible parts of plants that are consumed by humans or other animals as food. This original meaning is still commonly used, and is applied to plants collectively to refer to all edible plant matter, including edible flower, flo ...
in Chinese cuisine; they have a floral, yet
onion An onion (''Allium cepa'' , from Latin ), also known as the bulb onion or common onion, is a vegetable that is the most widely cultivated species of the genus '' Allium''. The shallot is a botanical variety of the onion which was classifie ...
-like flavor, attributed to volatile
organosulfur Organosulfur chemistry is the study of the properties and synthesis of organosulfur compounds, which are organic compounds that contain sulfur. They are often associated with foul odors, but many of the sweetest compounds known are organosulfur der ...
compounds. Plants with red young leaves are considered of better flavour than those where the young leaves are green.Plants for a Future
''Toona sinensis''
/ref>Oriental Vegetable Seeds

/ref> In China and Southeast Asian countries such as Malaysia, the young leaves of ''Toona sinensis'' or commonly known as Chinese Mahogany is used to make Toona paste, which is used as a condiment to serve with plain rice porridge as breakfast and simple meals, or to enhance the flavour of a dish or soup. Common dishes made with Toona paste are Chinese Mahogany fried rice, Chinese Mahogany beancurd, and Chinese Mahogany mushroom soup. The leaves contain Vitamin E, and high amounts of iron, calcium and
chlorophyll Chlorophyll is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words (, "pale green") and (, "leaf"). Chlorophyll allows plants to absorb energy ...
.


Lumber and landscape use

The
timber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...
is hard and reddish; it is valuable, used for furniture, musical instruments such as guitars, and in woodcarving. This is a common (and substantially cheaper) replacement for ''Swietenia'' or "true mahogany", which is now commercially restricted from being sourced natively. Outside Asia, ''T. sinensis'' is valued more as a large
ornamental tree Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...
for its haggard aspect. It is by far the most cold-tolerant species in the Meliaceae and the only member of the family that can be cultivated successfully in northern
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
.


Medicine

According to
Traditional Chinese Medicine Traditional Chinese medicine (TCM) is an alternative medicine, alternative medical practice drawn from traditional medicine in China. A large share of its claims are pseudoscientific, with the majority of treatments having no robust evidence ...
(TCM), the leaves of ''T. sinensis'' are beneficial for digestion and cough problems, and can help to stop bleeding. Recent researches find out that
polysaccharide Polysaccharides (), or polycarbohydrates, are the most abundant carbohydrates found in food. They are long-chain polymeric carbohydrates composed of monosaccharide units bound together by glycosidic linkages. This carbohydrate can react with wat ...
s contained in ''T. sinensis'' leaves can protect liver cells in high-fat or high-carbohydrate diets.
Quercetin Quercetin is a plant flavonol from the flavonoid group of polyphenols. It is found in many fruits, vegetables, leaves, seeds, and grains; capers, red onions, and kale are common foods containing appreciable amounts of it. It has a bitter flavor ...
extracted from leaves is found a natural antioxidant, and can act as a cancer preventive.


Chemical constituents

The leaves of ''T. sinensis'' has a relative high content of
nitrite The nitrite polyatomic ion, ion has the chemical formula . Nitrite (mostly sodium nitrite) is widely used throughout chemical and pharmaceutical industries. The nitrite anion is a pervasive intermediate in the nitrogen cycle in nature. The name ...
, which is about 157 to 160 mg/kg. However, after boiling with water, the amount of nitrite left is only about 7 mg/kg, which is safe to eat. Because the content of nitrite will increase as time passes, ''T. sinensis'' leaves are not suitable for long time storage. The
quercetin Quercetin is a plant flavonol from the flavonoid group of polyphenols. It is found in many fruits, vegetables, leaves, seeds, and grains; capers, red onions, and kale are common foods containing appreciable amounts of it. It has a bitter flavor ...
contained in ''T. sinensis'' leaves can induce
cell cycle The cell cycle, or cell-division cycle, is the sequential series of events that take place in a cell (biology), cell that causes it to divide into two daughter cells. These events include the growth of the cell, duplication of its DNA (DNA re ...
arrest and lead to
apoptosis Apoptosis (from ) is a form of programmed cell death that occurs in multicellular organisms and in some eukaryotic, single-celled microorganisms such as yeast. Biochemistry, Biochemical events lead to characteristic cell changes (Morphology (biol ...
of cells.


Culture

In
Chinese literature The history of Chinese literature extends thousands of years, and begins with the earliest recorded inscriptions, court archives, building to the major works of philosophy and history written during the Axial Age. The Han dynasty, Han (202  ...
, ''Toona sinensis'' is often used symbolically, with a mature tree representing a father. This manifests itself occasionally when expressing best wishes to a friend's father and mother in a letter, where one can write "wishing your ''Toona sinensis'' and
daylily A daylily, day lily or ditch-lily is a flowering plant in the genus ''Hemerocallis'' , a member of the family Asphodelaceae, subfamily Hemerocallidoideae, native to Asia. Despite the common name, it is not taxonomically classified in the lily gen ...
are strong and happy" (), with ''Toona sinensis'' metaphorically referring to the father and daylily to the mother.


References

{{Taxonbar, from=Q855853 sinensis Leaf vegetables Trees of China Flora of tropical Asia