''Tlatolophus'' (meaning "word crest") is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

hadrosaurid Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...

dinosaur belonging to the tribe

Parasaurolophini Lambeosauridae /ˌlæmbiəˈsɔːraɪniː/ (meaning 'lambe's lizards') is an extinct group of crested hadrosauroid dinosaurs. Description Size Uncertainty surrounds the size of lambeosaurs from the European continent. Hadrosaurs found there, ...

. The only species is the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

, ''Tlatolophus galorum''.

Discovery and naming

The

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

, specimen CIC/P/147, was first discovered as a semi-articulated tail in 2005 in sediments of the

Cerro del Pueblo Formation The Cerro del Pueblo Formation is a geological Formation (geology), formation in Saltillo, capital city of the state of Coahuila, Mexico, whose strata date back to the latest Campanian of the Late Cretaceous, just before the Campanian-Maastric ...

Coahuila Coahuila, formally Coahuila de Zaragoza, officially the Free and Sovereign State of Coahuila de Zaragoza, is one of the 31 states of Mexico. The largest city and State Capital is the city of Saltillo; the second largest is Torreón and the thi ...

Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...

. In 2013, the Mexican National Institute of Anthropology and History and the

National Autonomous University of Mexico The National Autonomous University of Mexico (, UNAM) is a public university, public research university in Mexico. It has several campuses in Mexico City, and many others in various locations across Mexico, as well as a presence in nine countri ...

launched a joint project to recover it. The holotype represents most complete

lambeosaurine Lambeosauridae /ˌlæmbiəˈsɔːraɪniː/ (meaning 'lambe's lizards') is an extinct group of crested hadrosauroid dinosaurs. Description Size Uncertainty surrounds the size of lambeosaurs from the European continent. Hadrosaurs found there, a ...

known from Mexico with an almost complete

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

, jaws, and additional parts of the

postcranial The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

skeleton including an articulated tail. It was named as the new genus and

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

''Tlatolophus galorum'' in 2021. The generic name comes from the

Náhuatl Nahuatl ( ; ), Aztec, or Mexicano is a language or, by some definitions, a group of languages of the Uto-Aztecan language family. Varieties of Nahuatl are spoken by about Nahuas, most of whom live mainly in Central Mexico and have smaller pop ...

word tlatolli, meaning "word", combined with

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

''lophos'' ("crest") due to the crest's resemblance to the

glyph A glyph ( ) is any kind of purposeful mark. In typography, a glyph is "the specific shape, design, or representation of a character". It is a particular graphical representation, in a particular typeface, of an element of written language. A ...

"word" in

Aztec The Aztecs ( ) were a Mesoamerican civilization that flourished in central Mexico in the Post-Classic stage, post-classic period from 1300 to 1521. The Aztec people included different Indigenous peoples of Mexico, ethnic groups of central ...

iconography, resembling an inverse comma. The

is named after the Garza and López families for their collaboration in collecting and preserving the specimen.

Description

The holotype specimen of ''Tlatolophus'', CIC/P/147, represents a large individual, around in length and weighing nearly . It preserves an almost complete skull, complete with lower jaws. CIC/P/147 only lacks the

surangular The surangular or suprangular is a jaw bone found in most land vertebrates, except mammals. Usually in the back of the jaw, on the upper edge, it is connected to all other jaw bones: dentary, angular bone, angular, splenial and articular. It is o ...

and quadrate bones. ''Tlatolophus'''s skull is relatively tall. Its

premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammals h ...

contacts the

nasal Nasal is an adjective referring to the nose, part of human or animal anatomy. It may also be shorthand for the following uses in combination: * With reference to the human nose: ** Nasal administration, a method of pharmaceutical drug delivery * ...

, but only along the nasal process. The

jugal The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy ...

orbital margins are wider than the infratemporal margin. The frontal platform is present, being dorsocaudally extended and overhanging the parietals. The

nasal cavity The nasal cavity is a large, air-filled space above and behind the nose in the middle of the face. The nasal septum divides the cavity into two cavities, also known as fossae. Each cavity is the continuation of one of the two nostrils. The nas ...

occupies the structure's entirety, including the caudal end of the supracranial crest which is comma-shaped, hence the

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...

. The

pterygoid bone The pterygoid is a paired bone forming part of the palate of many vertebrates, behind the palatine bone In anatomy, the palatine bones (; derived from the Latin ''palatum'') are two irregular bones of the facial skeleton in many animal specie ...

possesses a high and convex dorsal crest. CIC/P/147 also preserves partial postcranial elements. The scapular neck is relatively stout, and the iliac postacetabular process is long. The ischium possesses a large ischial foot, and its tip is cranioventrally projected. A partial, articulated tail is also preserved, the first remains to be discovered of this taxon.

Classification

''Tlatolophus'' was assigned to the tribe Parasaurolophini by Ramírez-Velasco ''et al.'' in 2021. The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below displays the results of their

phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...

References

{{Taxonbar, from=Q106805159 Lambeosaurinae Dinosaur genera Campanian dinosaurs Fossil taxa described in 2021 Dinosaurs of Mexico

Discovery and naming

Description

Classification

See also

References