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Timothy (''Phleum pratense'') is an abundant
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
grass Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family (biology), family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and spe ...
native to most of Europe except for the Mediterranean region. It is also known as timothy-grass, meadow cat's-tail or common cat's tail. It is a member of the genus '' Phleum'', consisting of about 15 species of annual and perennial grasses. It is probably named after Timothy Hanson, an American farmer and agriculturalist said to have introduced it from
New England New England is a region consisting of six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York (state), New York to the west and by the ...
to the southern states in the early 18th century. Upon his recommendation it became a major source of hay and cattle fodder to British
farmer A farmer is a person engaged in agriculture, raising living organisms for food or raw materials. The term usually applies to people who do some combination of raising field crops, orchards, vineyards, poultry, or other livestock. A farmer ...
s in the mid-18th century. Timothy can be confused with meadow foxtail (''Alopecurus pratensis'') or purple-stem cat's-tail (''Phleum phleoides'').


Description

Timothy grows to tall, with
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
up to long and broad. The leaves are hairless, rolled rather than folded, and the lower sheaths turn dark brown. It has no
stolon In biology, a stolon ( from Latin ''wikt:stolo, stolō'', genitive ''stolōnis'' – "branch"), also known as a runner, is a horizontal connection between parts of an organism. It may be part of the organism, or of its skeleton. Typically, animal ...
s or
rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...
s, and no auricles. The
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
head is long and broad, with densely packed spikelets. It flowers from June until September. The
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
are pink. The
ligule A ligule (from "strap", variant of ''lingula'', from ''lingua'' "tongue") is a thin outgrowth at the junction of leaf A leaf (: leaves) is a principal appendage of the plant stem, stem of a vascular plant, usually borne laterally above g ...
is short and blunt. It grows well in heavy soil, and is noted for its resistance to cold and drought, and thus ability to grow in dry upland or poor sandy soils. In pasture it tends to be overwhelmed by more competitive grasses. After cutting it grows slowly. File:Poaceae. - Phleum pratense.JPG, Close-up of flower head showing purple stamen (3 per floret) and feathery stigma (2 per floret) File:Timothy ligule.jpg, Ligule is short and blunt File:Illustration Phleum pratense0.jpg, Showing bulbous base and brown leaf sheaths File:Timothy seeds.jpg, Seeds


Subspecies

There are two
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
: * ''Phleum pratense'' subsp. ''pratense''. Larger, to tall. Widespread. Native to the Mediterranean. * ''Phleum pratense'' subsp. ''bertolonii''. Smaller, to tall. Calcareous grassland.


Cultivation and uses

Timothy was unintentionally introduced to North America by early settlers, and was first described in 1711 by John Hurd from plants growing in
New Hampshire New Hampshire ( ) is a U.S. state, state in the New England region of the Northeastern United States. It borders Massachusetts to the south, Vermont to the west, Maine and the Gulf of Maine to the east, and the Canadian province of Quebec t ...
. Hurd named the grass "hurd grass" but a farmer named Timothy Hanson began to promote cultivation of it as a hay about 1720, and the grass has been known by its present name since then. Timothy has now become naturalized throughout most of the US and Canada. It is commonly grown for
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
feed and, in particular, as hay for
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
s. It is relatively high in fiber, especially when cut late. It is considered a harsh, coarse grass little relished by livestock if cut earlier. It is considered part of the standard mix for grass hay and provides quality nutrition for horses. Timothy hay is a staple food for domestic pet rabbits,
guinea pig The guinea pig or domestic guinea pig (''Cavia porcellus''), also known as the cavy or domestic cavy ( ), is a species of rodent belonging to the genus ''Cavia'', family Caviidae. Animal fancy, Breeders tend to use the name "cavy" for the ani ...
s, chinchillas, and degus, often making up the bulk of their diet. Timothy hay is rich in long fiber and its abrasive texture helps to grind down the teeth, keeping both the teeth and jaw in good order. Some
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s use it as a food plant, e.g. the Essex skipper (''Thymelicus lineola'') and the marbled white (''Melanargia galathea''). It also grows in roadsides and abandoned fields but generally requires nutrient-rich soils. Plants persist through the winter. Dead, straw-colored flowering stems may persist, but only for a short time, and are recognized by the distinctive spike-like
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
.


Medical uses

Timothy grass pollen allergen is used to treat
hay fever Allergic rhinitis, of which the seasonal type is called hay fever, is a type of rhinitis, inflammation in the nose that occurs when the immune system overreacts to allergens in the air. It is classified as a Allergy, type I hypersensitivity re ...
.


Breeding objectives in timothy

Breeding programs for forage grasses and especially timothy have been focusing on the improvement of dry matter yield, resistance to disease, dry matter digestibility, and nutritional value, which depends on target species and environment. Due to high phenotypic and genetic heterogeneity in individual plants, and the polyploidy of many species, breeding programs for timothy are accompanied by some difficulties.


Confusion with other species

It is often confused with meadow foxtail (''Alopecurus pratensis''). Timothy flowers later, from June until August, whereas meadow foxtail flowers from April until June. The
spikelet A spikelet, in botany, describes the typical arrangement of the inflorescences of grasses, sedges and some other monocots. Each spikelet has one or more florets. The spikelets are further grouped into panicles or spikes. The part of the sp ...
s of timothy are twin hornlike projections arranged in cylindrical
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
s, whereas foxtail has a soft, single awn. Purple-stem cat's-tail (''Phleum phleoides'') prefers lighter soils and grows on chalk downland. Mountain timothy (''Phleum alpinum'') grows above . A "wild Timothy" was found to grow in Yosemite at the time of its discovery but may have been a foxtail.Scribner, F. Lampson, "Grasses of Mountain Meadows and Deer Parks" in Timothy canary grass (''Phalaris angusta''), another species with a similar cylindrical panicle, is toxic to livestock. File:Timothy awns.jpg, Close-up of flower head showing hornlike spikelets File:Alopecurus pratensis1.JPG, Meadow foxtail (''
Alopecurus pratensis ''Alopecurus pratensis'', known as the meadow foxtail or the field meadow foxtail, is a Perennial plant, perennial grass belonging to the grass family (Poaceae). It is native to Europe and Asia. This common plant is found on grasslands, especia ...
'') spikelet


References


External links


Flora Europaea: ''Phleum pratense''

Timothy - US Department of Agriculture
{{Taxonbar, from=Q256508 Pooideae Bunchgrasses of Europe Flora of Eastern Europe Flora of Middle Europe Flora of Northern Europe Flora of Estonia Forages Medicinal plants of Europe Plants described in 1753 Grasses of Lebanon