Tarantula Hawk
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A tarantula hawk is a spider wasp (Pompilidae) that preys on
tarantula Tarantulas comprise a group of large and often hairy spiders of the family Theraphosidae. , 1,100 species have been identified, with 166 genera. The term "tarantula" is usually used to describe members of the family Theraphosidae, although ...
s. Tarantula hawks belong to any of the many species in the genera '' Pepsis'' and '' Hemipepsis.'' They are some of the largest
parasitoid wasp Parasitoid wasps are a large group of hymenopteran Superfamily (zoology), superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, ...
s, using their sting to paralyze their prey before dragging it into a brood nest as living food; a single egg is laid on the prey, hatching to a larva, which then eats the still-living host. They are found on all continents other than
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
.


Description

These wasps grow up to long, making them among the largest of wasps, and have blue-black bodies and bright, rust-colored wings (other species have black wings with blue highlights). The vivid coloration found on their bodies, and especially wings, is aposematic, advertising to potential predators the wasps' ability to deliver a powerful sting. Their long legs have hooked claws for grappling with their victims. The stinger of a female '' Pepsis grossa'' can be up to long, and the powerful sting is considered one of the most painful insect stings in the world.


Behavior

The female tarantula hawk wasp stings a tarantula between the legs, paralyzing it, and then drags the prey to a specially prepared burrow, where a single egg is laid on the spider's abdomen, and the burrow entrance is covered. Sex of offspring is determined by fertilization; fertilized eggs produce females, while unfertilized eggs produce males. When the wasp
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
hatches, it creates a small hole in the spider's
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
, then enters and feeds voraciously, avoiding vital organs for as long as possible to keep the spider alive. After several weeks, the larva
pupa A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages th ...
tes. Finally, the wasp becomes an adult and emerges from the spider's abdomen to continue the life cycle. Adult tarantula hawks are nectarivorous. While the wasps tend to be most active in the daytime in summer, they tend to avoid high temperatures. The male tarantula hawk does not hunt. Both males and females feed on the flowers of milkweeds, western soapberry trees, or mesquite trees. Male tarantula hawks have been observed practicing a behavior called hill-topping, in which they sit atop tall plants and watch for passing females ready to reproduce. The males can become resident defenders of the favorable reproduction spots for hours into the afternoon. Females are not very aggressive, in that they are hesitant to sting, but the sting is extraordinarily painful.


Distribution

Worldwide distribution of tarantula hawks includes areas from
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
to
Southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
, Africa, Australia, and the Americas, with the genus '' Pepsis'' entirely restricted to the New World. In the latter, ''Pepsis'' species have been observed from as far north as
Logan, Utah Logan is a city in Cache County, Utah, United States. The 2020 United States Census, 2020 census recorded the population at 52,778. Logan is the county seat of Cache County and the principal city of the Logan metropolitan area, which includes Ca ...
, and south as far as
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
, with at least 250 species living in South America. Eighteen species of ''Pepsis'' and three species of '' Hemipepsis'' are found in the United States, primarily in the deserts of the Southwestern United States, with '' Pepsis grossa'' (formerly ''P. formosa'') and '' Pepsis thisbe'' being common. The two species are difficult to distinguish, but most ''P. grossa'' wasps have metallic blue bodies and reddish antennae, which separate them from ''P. thisbe''. Both species have bright orange wings that become transparent near the tip.


Sting

Tarantula hawk wasps are relatively docile and rarely sting without provocation, but the sting—particularly that of ''P. grossa''—is among the most painful of all insects, though the intense pain only lasts about five minutes. One researcher described the pain as "...immediate, excruciating, unrelenting pain that simply shuts down one's ability to do anything, except scream. Mental discipline simply does not work in these situations." In terms of scale, the wasp's sting is rated near the top of the Schmidt sting pain index, second only to that of the bullet ant, and is described by Schmidt as "blinding, fierce andshockingly electric". Because of their extremely large stingers, very few animals are able to eat them; one of the few that can is the roadrunner. Many predatory animals avoid these wasps, and many different insects
mimic In evolutionary biology, mimicry is an evolved resemblance between an organism and another object, often an organism of another species. Mimicry may evolve between different species, or between individuals of the same species. In the simples ...
them, including various other wasps and bees ( Müllerian mimics), as well as
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
s,
flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwin ...
(e.g., mydas flies), and
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s (e.g., '' Tragidion'') ( Batesian mimics). Aside from the possibility of triggering an allergic reaction, the sting is not dangerous and does not require medical attention. Local redness appears in most cases after the pain, and lasts for up to a week.


State insect of New Mexico

The U.S. state of
New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
chose a species of tarantula hawk (specifically, ''P. formosa'', now known as ''P. grossa'') in 1989 to become its official state insect. Its selection was prompted by a group of elementary-school children from Edgewood doing research on states that had adopted state insects. They selected three insects as candidates and mailed ballots to all schools for a statewide election. The winner was the tarantula hawk wasp.


References


External links

{{Wikispecies, Pepsis
'"Instantaneous, Electrifying, Excruciating Pain"
book excerpt by entomologist Justin O. Schmidt Pepsinae Aposematic species