''Steropodon'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of prehistoric

monotreme Monotremes () are prototherian mammals of the order Monotremata. They are one of the three groups of living mammals, along with placentals ( Eutheria), and marsupials (Metatheria). Monotremes are typified by structural differences in their brai ...

, or egg-laying

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...

. It contains a single species, ''Steropodon galmani'', that lived about 105 to 93.3 million years ago (mya) in the

Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

period. It is one of the oldest monotremes discovered, and is one of the oldest

Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...

n mammal discoveries.

Taxonomy

The dentition of ''Steropodon'' is somewhat similar to that of

theria Theria (; Greek: , wild beast) is a subclass of mammals amongst the Theriiformes. Theria includes the eutherians (including the placental mammals) and the metatherians (including the marsupials) but excludes the egg-laying monotremes. ...

ns—the placentals and the

marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in ...

s—specifically the presence of the

tribosphenic molar The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...

tooth which was thought to be exclusive to therians since the Cretaceous. This, along with the tribosphenic molar discoveries of monotreme-relatives '' Ausktribosphenos'' and ''

Ambondro mahabo ''Ambondro mahabo'' is a mammal from the Middle Jurassic (Bathonian) Isalo III Formation (about 167 million years ago) of Madagascar. The only described species of the genus ''Ambondro'', it is known from a fragmentary lower jaw with three teeth ...

'' of which the latter evolved in the Jurassic, led to the conclusion that the molar evolved independently in the two lineages. This inspired the creation of the subclasses Australosphenida—the monotremes and extinct relatives—and

Tribosphenida Tribosphenida is a group (infralegion) of mammals that includes the ancestor of ''Hypomylos'', Aegialodontia and Theria (the last common ancestor of marsupials and placentals plus all of its descendants). Its current definition is more or less sy ...

—placentals and marsupials. However, given this classification is based only on jaw and lower-tooth remains, insufficient evidence may exist to definitively make this conclusion. It may be only described member of the family Steropodontidae. Another proposed member is ''

Teinolophos ''Teinolophos'' is a prehistoric species of monotreme, or egg-laying mammal, from the Teinolophidae. It is known from four specimens, each consisting of a partial lower jawbone collected from the Wonthaggi Formation at Flat Rocks, Victoria, Aus ...

'' that was moved into its own family, Teinolophidae, by Flannery ''et al.'' (2022). Edentulous partial

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...

from the Finch Clay facies of the

Griman Creek Formation The Griman Creek Formation is a geological formation in northern New South Wales and southern Queensland, Australia whose strata date back to the Albian-Cenomanian of the Early-Late Cretaceous.Bell et al., 2019 It is most notable being a major so ...

was attributed to undescribed steropodontid by Musser (2013). On other side, Flannery ''et al.'' assumed that this mandible may belong to a new genus and species of

stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...

ornithorhynchid The Ornithorhynchidae are one of the two extant families in the order Monotremata, and contain the platypus and its extinct relatives. The other family is the Tachyglossidae, or echidnas. Within the Ornithorhynchidae are the genera '' Monotrem ...

. The specific epithet is derived from the surname of the jaw's collectors, when combined with the genus name ''Steropodon'' is intended to translate as "Galman's lightning tooth".

Description

''Steropodon'' is known only from a single

opal Opal is a hydrated amorphous form of silica (SiO2·''n''H2O); its water content may range from 3 to 21% by weight, but is usually between 6 and 10%. Due to its amorphous property, it is classified as a mineraloid, unlike crystalline form ...

ised jaw with three molars, discovered at the

, Lightning Ridge,

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

, Australia, by brothers David and Alan Galman. It was a large

for the

Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...

, being long. The lower molars are in length, with a width of . A length of is more typical for Mesozoic mammals. Woodburne (2003, p. 212) reports that the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

is a right mandible named AM F66763. The preserved molars are m1–m3. Examination of the jaw fragment revealed a mandibular canal, which has been proposed to indicate the presence of a bill, similar to those of the extinct species ''

Obdurodon dicksoni ''Obdurodon'' is a genus of extinct platypus-like Australian monotreme which lived from the Late Oligocene to the Late Miocene. Three species have been described in the genus, the type species ''Obdurodon insignis'', plus ''Obdurodon dicksoni'' ...

'' and the modern platypus ''

Ornithorhynchus anatinus The platypus (''Ornithorhynchus anatinus''), sometimes referred to as the duck-billed platypus, is a semiaquatic, egg-laying mammal endemic to eastern Australia, including Tasmania. The platypus is the sole living representative or monotypi ...

''.

References

External links

Steropodon
at Fossilworks * * {{Taxonbar, from=Q131338 Prehistoric monotremes Prehistoric mammals of Australia Cretaceous mammals Prehistoric mammal genera Fossil taxa described in 1985 Taxa named by Mike Archer (paleontologist) Taxa named by Tim Flannery Taxa named by Ralph Molnar Cretaceous animals of Australia

Taxonomy

Description

See also

References

Further reading

External links