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The sika deer (''Cervus nippon''), also known as the Northern spotted deer or the Japanese deer, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...
native to much of East Asia and introduced to other parts of the world. Previously found from northern
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making ...
in the south to the
Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admin ...
in the north, it is now uncommon except in
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...
, where the species is overabundant.


Etymology

Its name comes from , the Japanese word for "deer". In Japan, the species is known as the . In Chinese, it is known as .


Taxonomy

The sika deer is a member of the genus '' Cervus'', a group of deer also known as the "true deer". Formerly, sika were grouped together in this genus with nine other species. Now, only the sika and red deer remain, the latter being divided into three separate species: European red deer,
central Asian red deer The Central Asian red deer (''Cervus hanglu''), also known as the Tarim red deer is a deer species native to Central Asia, where it used to be widely distributed, but is scattered today with small population units in several countries. It has be ...
, and American elk (though this remains controversial). Recent DNA evidence indicates these deer are not as closely related as previously thought, resulting in the creation of new species and genera. The genera '' Rucervus'', '' Rusa'', and '' Przewalskium'' are where most of the former ''Cervus'' species now belong. The ancestor of all ''Cervus'' species probably originated in central Asia and resembled sika deer. All ''Cervus'' species can crossbreed and produce hybrids in areas where they coexist (for example, introduced sika hybridize with native red deer in the
Scottish Highlands The Highlands ( sco, the Hielands; gd, a’ Ghàidhealtachd , 'the place of the Gaels') is a historical region of Scotland. Culturally, the Highlands and the Lowlands diverged from the Late Middle Ages into the modern period, when Lowland S ...
, where this is a serious threat to the gene pool of the red deer population).


Subspecies

Serious
genetic pollution Genetic pollution is a controversial term for uncontrolled gene flow into wild populations. It is defined as "the dispersal of contaminated altered genes from genetically engineered organisms to natural organisms, esp. by cross-pollination", but ...
has occurred in many populations, especially in China, so the status of many
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
remains unclear. The status of ''C. n. hortulorum'' is particularly uncertain and might in fact be of mixed origin, hence it is not listed here. *'' C. n. aplodontus'', northern
Honshu , historically called , is the largest and most populous island of Japan. It is located south of Hokkaidō across the Tsugaru Strait, north of Shikoku across the Inland Sea, and northeast of Kyūshū across the Kanmon Straits. The island s ...
*'' C. n. grassianus'',
Shanxi Shanxi (; ; formerly romanised as Shansi) is a landlocked province of the People's Republic of China and is part of the North China region. The capital and largest city of the province is Taiyuan, while its next most populated prefecture-leve ...
, China *'' C. n. keramae'', Kerama Islands of the
Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Taiwan: the Ōsumi, Tokara, Amami, Okinawa, and Sakishima Islands (further divided into the Miyako and Yaeyama Islands), with Yon ...
, Japan *'' C. n. kopschi'', southern China *'' C. n. mandarinus'', northern and northeastern China *'' C. n. mantchuricus'', northeastern China,
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic ...
, and
Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admin ...
*'' C. n. nippon'', southern
Honshu , historically called , is the largest and most populous island of Japan. It is located south of Hokkaidō across the Tsugaru Strait, north of Shikoku across the Inland Sea, and northeast of Kyūshū across the Kanmon Straits. The island s ...
,
Shikoku is the smallest of the four main islands of Japan. It is long and between wide. It has a population of 3.8 million (, 3.1%). It is south of Honshu and northeast of Kyushu. Shikoku's ancient names include ''Iyo-no-futana-shima'' (), '' ...
, and
Kyushu is the third-largest island of Japan's five main islands and the most southerly of the four largest islands ( i.e. excluding Okinawa). In the past, it has been known as , and . The historical regional name referred to Kyushu and its surround ...
*'' C. n. pseudaxis'', northern
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making ...
*'' C. n. pulchellus'',
Tsushima Island is an island of the Japanese archipelago situated in-between the Tsushima Strait and Korea Strait, approximately halfway between Kyushu and the Korean Peninsula. The main island of Tsushima, once a single island, was divided into two in 167 ...
*'' C. n. sichuanicus'', western China *'' C. n. sintikuensis'',
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the no ...
*'' C. n. soloensis'', Southern
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
(anciently introduced to
Jolo island Jolo ( tsg, Sūg) is a volcanic island in the southwest Philippines and the primary island of the province of Sulu, on which the capital of the same name is situated. It is located in the Sulu Archipelago, between Borneo and Mindanao, and has ...
; of unknown subspecies origin, probably extinct) *'' C. n. taiouanus'',
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the no ...
*'' C. n. yakushimae'',
Yakushima is one of the Ōsumi Islands in Kagoshima Prefecture, Japan. The island, in area, has a population of 13,178. Access to the island is by hydrofoil ferry (7 or 8 times a day from Kagoshima, depending on the season), slow car ferry (once or twic ...
, Japan *'' C. n. yesoensis'',
Hokkaido is Japan, Japan's Japanese archipelago, second largest island and comprises the largest and northernmost Prefectures of Japan, prefecture, making up its own List of regions of Japan, region. The Tsugaru Strait separates Hokkaidō from Honshu; th ...
, Japan


Description

The sika deer is one of the few deer species that does not lose its spots upon reaching maturity. Spot patterns vary with region. The mainland subspecies have larger and more obvious spots, in contrast to the
Taiwanese Taiwanese may refer to: * Taiwanese language, another name for Taiwanese Hokkien * Something from or related to Taiwan (Formosa) * Taiwanese aborigines, the indigenous people of Taiwan * Han Taiwanese, the Han people of Taiwan * Taiwanese people, r ...
and Japanese subspecies, whose spots are nearly invisible. Many introduced populations are from Japan, so they also lack significant spots. The color of the
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...
ranges from mahogany to black, and white individuals are also known. During winter, the coat becomes darker and shaggier and the spots less prominent, and a mane forms on the back of the males' necks. They are medium-sized
herbivores A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, though they show notable size variation across their several subspecies and considerable
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
, with males invariably much larger than females. They can vary from tall at the shoulder and from in head-and-body length. The tail measures about long. The largest subspecies is the Manchurian sika deer (''C. n. mantchuricus''), in which males commonly weigh about and females weigh , with large stags scaling up to , although there had been records of
Yezo sika deer The Yezo sika deer (''Cervus nippon yesoensis'', ja, エゾシカ / 蝦夷鹿, yezoshika, link=no, Ainu: ユク ''yuk'') is one of the many subspecies of the sika deer. The sika that inhabit the island of Hokkaido are indigenous, although it i ...
bulls weighing up to .『エゾシカは森の幸 人・森・シカの共生』p.63 On the other end of the size spectrum, in the Japanese sika deer (''C. n. nippon''), males weigh and females weigh . All sikas are compact and dainty-legged, with short, trim, wedge-shaped heads and a boisterous disposition. When alarmed, they often display a distinctive flared rump, much like the American elk. Sika stags have stout, upright
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on ...
s with an extra buttress up from the brow
tine Tine may refer to: *Tine (structural), a 'prong' on a fork or similar implement, or any similar structure *Tine (company), the biggest dairy producer in Norway *Tine (film), ''Tine'' (film), a 1964 Danish film *Tine, Iran, a village in Mazandaran P ...
and a very thick wall. A forward-facing intermediate tine breaks the line to the top, which is usually forked. Occasionally, sika antlers develop some palmation (flat areas). Females carry a pair of distinctive black bumps on the forehead. Antlers can range from to more than , depending on the subspecies. Stags also have distinctive manes during their mating period ( rut). File:Cervus nippon Kitz Juni 2012 Wildpark Alte Fasanerie Klein-Auheim.JPG, Fawn at the Wildpark Alte Fasanerie in Klein-Auheim File:Hokkaido Sika Deer.jpg, In Shiretoko Peninsula,
Hokkaido is Japan, Japan's Japanese archipelago, second largest island and comprises the largest and northernmost Prefectures of Japan, prefecture, making up its own List of regions of Japan, region. The Tsugaru Strait separates Hokkaidō from Honshu; th ...
, Japan Image:Cervus nippon antlers - Finnish Museum of Natural History - DSC04525.JPG, The skull of stag displayed in the
Finnish Museum of Natural History The Finnish Museum of Natural History ( fi, Luonnontieteellinen keskusmuseo, sv, Naturhistoriska centralmuseet), established in 1988, is a research institution under the University of Helsinki in Finland, based in Helsinki, Finland. It is a natur ...
,
Helsinki Helsinki ( or ; ; sv, Helsingfors, ) is the Capital city, capital, primate city, primate, and List of cities and towns in Finland, most populous city of Finland. Located on the shore of the Gulf of Finland, it is the seat of the region of U ...
, Finland File:Cervus nippon yakushimae - 20160710.webm, Sika deer standing up. File:Sika deer in Nara 05.jpg, Young male in
Nara The National Archives and Records Administration (NARA) is an " independent federal agency of the United States government within the executive branch", charged with the preservation and documentation of government and historical records. It ...


Behavior

Sika deer can be active throughout the day, though in areas with heavy human disturbance, they tend to be
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
. Seasonal migration is known to occur in mountainous areas, such as Japan, with winter ranges being up to lower in elevation than summer ranges. Lifestyles vary between individuals, with some occurring alone while others are found in single-sex groups. Large herds gather in autumn and winter. Males spend most years alone occasionally forming herds together. Females with fawns only form herds during birthing season. The sika deer is a highly vocal species, with over 10 individual sounds, ranging from soft whistles to loud screams. Sika males are territorial and keep
harem Harem ( Persian: حرمسرا ''haramsarā'', ar, حَرِيمٌ ''ḥarīm'', "a sacred inviolable place; harem; female members of the family") refers to domestic spaces that are reserved for the women of the house in a Muslim family. A har ...
s of females during their rut, which peaks from early September through November, but may last well into the winter. Territory size varies with habitat type and size of the buck; strong, prime bucks may hold up to . Territories are marked by a series of shallow pits or "scrapes", which is digging holes (up to 1.6 m in wide and 0.3 m in deep) with forefeet or antlers, into which the males urinate and from which emanates a strong,
musk Musk ( Persian: مشک, ''Mushk'') is a class of aromatic substances commonly used as base notes in perfumery. They include glandular secretions from animals such as the musk deer, numerous plants emitting similar fragrances, and artificial sub ...
y odor. Fights between rival males for territorial disputes, which occur by using hooves and antlers, are sometimes fierce and long and may even be fatal. The
gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once ...
lasts for seven months. Hinds (does) give birth to a single fawn, weighing , which is nursed for up to ten months. The mother hides her fawn in thick undergrowth immediately after giving birth, and the fawn stays very quiet and still while it waits until the mother returns to nurse it. The fawn becomes independent 10 to 12 months after birth, and attains sexual maturity at 16 to 18 months in both sexes. The average lifespan is 15 to 18 years in captivity, although one case is recorded as living 25 years and 5 months. The sika deer may interbreed with the
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of ...
, the closest relative; hybrid descendants may have adaptive advantages over purebred relatives. In
Nara Prefecture is a prefecture of Japan located in the Kansai region of Honshu. Nara Prefecture has a population of 1,321,805 and has a geographic area of . Nara Prefecture borders Kyoto Prefecture to the north, Osaka Prefecture to the northwest, Wakaya ...
, Japan, the deer are also known as "bowing deer", as they bow their heads before being fed special . However, deer bow heads to signal that they are about to
headbutt A headbutt is a targeted strike with the head, typically (when intentional) involving the use of robust parts of the headbutter's cranium as the area of impact. The most effective headbutts strike the most sensitive areas of an opponent, such as ...
. Therefore, when a human "bows" to a deer, the deer may take it as a challenge, and will assume the same stance before charging and attempting to headbutt the person. Deer headbutt both for play and to assert dominance, as do goats. Sika deer are found throughout the city of Nara and its many parks and temples like
Tōdai-ji is a Buddhist temple complex that was once one of the powerful Seven Great Temples, located in the city of Nara, Japan. Though it was originally founded in the year 738 CE, Tōdai-ji was not opened until the year 752 CE. The temple has undergo ...
, as they are considered to be the messengers of the
Shinto Shinto () is a religion from Japan. Classified as an East Asian religion by scholars of religion, its practitioners often regard it as Japan's indigenous religion and as a nature religion. Scholars sometimes call its practitioners ''Shint ...
gods.


Habitat

Sika deer are found in the
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
and
subtropical The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° north a ...
forests of eastern Asia, preferring areas with dense understory, and where snowfall does not exceed . They tend to forage in patchy clearings of forests. Introduced populations are found in areas with similar habitats to their native ranges, including Western and Central Europe, Eastern United States, and
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
.


Population

Sika deer inhabit temperate and subtropical woodlands, often in areas suitable for farming and other human exploitation. Their range encompasses some of the most densely populated areas in the world, where forests were cleared hundreds of years ago. Their population status varies significantly in different countries. Although the species as a whole is thriving, it is endangered and extinct in many areas. Japan has by far the largest native sika population in the world. Though the exact population is uncertain, it is likely to be in the hundred thousand range and is still increasing, mainly due to recent conservation efforts and the extinction of its main predator, the Japanese wolf (''Canis lupus hodophilax''), over a century ago. Without its main predator, the population of sika exploded and it is now overpopulated in many areas, posing a threat to both forests and farmlands. Efforts are now being made to control its population instead of conserving it. None of its subspecies is endangered except the Kerama deer (''C. n. keramae'') on the tiny Kerama Islands. In 2015,
Japanese Ministry of the Environment The is a Cabinet-level ministry of the government of Japan responsible for global environmental conservation, pollution control, and nature conservation. The ministry was formed in 2001 from the sub-cabinet level Environmental Agency establish ...
estimated the population at 3,080,000 in Japan, including
Hokkaido is Japan, Japan's Japanese archipelago, second largest island and comprises the largest and northernmost Prefectures of Japan, prefecture, making up its own List of regions of Japan, region. The Tsugaru Strait separates Hokkaidō from Honshu; th ...
. China used to have the largest population of sika, but thousands of years of hunting and habitat loss have reduced the population to less than 1,000. Of the five subspecies in China, the North China sika deer (''C. n. mandarinus'') is believed to be extinct in the wild since the 1930s; the Shanxi sika deer (''C. n. grassianus'') has not been seen in the wild since the 1980s and is also believed to be extinct in the wild. The status of Manchurian sika deer in China is unclear, though it is also believed to be extinct, and the sightings there are actually feral populations. The South China sika deer (''C. n. kopschi'') and Sichuan sika deer (''C. n. sichuanicus'') are the only subspecies known to remain in the wild in China. The former exists in fragmented populations of around 300 in southeast China, while the latter is found in a single population of over 400. The feral population is likely to be much higher than the wild, though most of them are descended from domesticated sikas of mixed subspecies. All of the subspecies are present in captivity, but a lack of suitable habitats and government efforts prevent their reintroduction. The Formosan sika deer (''C. n. taioanus'') has been extinct in the wild for almost two decades before individuals from zoos were introduced to
Kenting National Park Kenting National Park (), commonly known as Kenting (), is a national park located on the Hengchun Peninsula of Pingtung County, Taiwan, covering Hengchun, Checheng, and Manzhou Townships. Established on 1 January 1984, it is Taiwan's oldest ...
; the population now numbers 200. Reintroduction programs are also under way in
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making ...
, where the Vietnamese sika deer (''C. n. pseudaxis'') is
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
or nearly so.
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
has a relatively large and stable population of 8,500–9,000 individuals of the Manchurian subspecies, but this is limited to a small area in
Primorsky Krai Primorsky Krai (russian: Приморский край, r=Primorsky kray, p=prʲɪˈmorskʲɪj kraj), informally known as Primorye (, ), is a federal subject (a krai) of Russia, located in the Far East region of the country and is a part of t ...
. Small populations might exist in
North Korea North Korea, officially the Democratic People's Republic of Korea (DPRK), is a country in East Asia. It constitutes the northern half of the Korean Peninsula and shares borders with China and Russia to the north, at the Yalu (Amnok) and T ...
, but the political situation makes investigation impossible. The species is extinct in
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
, with no plans for reintroduction. File:Sikadeer.jpg, Outside of a store on the island of
Miyajima may refer to: Places * Miyajima, another name for the Japanese island Itsukushima * Miyajima, Hiroshima, a former town on this island, merged into Hatsukaichi, Hiroshima in 2005 * Itsukushima Shrine is a Shinto shrine on the island of Itsukus ...
File:Miyajima Deer Sep08.jpg, Tame deer wandering the streets of the town of
Miyajima may refer to: Places * Miyajima, another name for the Japanese island Itsukushima * Miyajima, Hiroshima, a former town on this island, merged into Hatsukaichi, Hiroshima in 2005 * Itsukushima Shrine is a Shinto shrine on the island of Itsukus ...
, Japan


Introduced populations

Sika deer have been introduced into a number of other countries, including
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, an ...
,
Latvia Latvia ( or ; lv, Latvija ; ltg, Latveja; liv, Leţmō), officially the Republic of Latvia ( lv, Latvijas Republika, links=no, ltg, Latvejas Republika, links=no, liv, Leţmō Vabāmō, links=no), is a country in the Baltic region of ...
,
Lithuania Lithuania (; lt, Lietuva ), officially the Republic of Lithuania ( lt, Lietuvos Respublika, links=no ), is a country in the Baltic region of Europe. It is one of three Baltic states and lies on the eastern shore of the Baltic Sea. Lithuania ...
,
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
,
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to ...
,
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of Denmark , establish ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwee ...
,
Ireland Ireland ( ; ga, Éire ; Ulster-Scots: ) is an island in the North Atlantic Ocean, in north-western Europe. It is separated from Great Britain to its east by the North Channel, the Irish Sea, and St George's Channel. Ireland is the s ...
,
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
,
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of ...
,
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
,
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
,
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
,
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
, the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
(
Jolo Island Jolo ( tsg, Sūg) is a volcanic island in the southwest Philippines and the primary island of the province of Sulu, on which the capital of the same name is situated. It is located in the Sulu Archipelago, between Borneo and Mindanao, and has ...
),
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic countries, Nordic c ...
,
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bot ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...
, the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and ...
, and the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
(
Delaware Delaware ( ) is a state in the Mid-Atlantic region of the United States, bordering Maryland to its south and west; Pennsylvania to its north; and New Jersey and the Atlantic Ocean to its east. The state takes its name from the adjacent ...
,
Kansas Kansas () is a U.S. state, state in the Midwestern United States, Midwestern United States. Its Capital city, capital is Topeka, Kansas, Topeka, and its largest city is Wichita, Kansas, Wichita. Kansas is a landlocked state bordered by Nebras ...
,
Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean t ...
,
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Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...
). In many cases, they were originally introduced as ornamental animals in parklands, but have established themselves in the wild. On Spieden Island in the
San Juan Islands The San Juan Islands are an archipelago in the Pacific Northwest of the United States between the U.S. state of Washington and Vancouver Island, British Columbia, Canada. The San Juan Islands are part of Washington state, and form the core ...
of Washington, they were introduced as a game animal. In the UK and Ireland, several distinct feral populations now exist. Some of these are in isolated areas, for example on the island of
Lundy Lundy is an English island in the Bristol Channel. It was a micronation from 1925–1969. It forms part of the district of Torridge in the county of Devon. About long and wide, Lundy has had a long and turbulent history, frequently chang ...
, but others are contiguous with populations of the native
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of ...
. Since the two species sometimes hybridize, a serious conservation concern exists. In research which rated the negative impact of introduced mammals in Europe, the sika deer was found to be among the most damaging to the environment and economy, along with the brown rat and
muskrat The muskrat (''Ondatra zibethicus'') is a medium-sized semiaquatic rodent native to North America and an introduced species in parts of Europe, Asia, and South America. The muskrat is found in wetlands over a wide range of climates and habita ...
. In the 1900s,
King Edward VII Edward VII (Albert Edward; 9 November 1841 – 6 May 1910) was King of the United Kingdom of Great Britain and Ireland and Emperor of India, from 22 January 1901 until his death in 1910. The second child and eldest son of Queen Victoria an ...
presented a pair of sika deer to John, the second Baron Montagu of Beaulieu. This pair escaped into Sowley Wood and were the basis of the sika to be found in the
New Forest The New Forest is one of the largest remaining tracts of unenclosed pasture land, heathland and forest in Southern England, covering southwest Hampshire and southeast Wiltshire. It was proclaimed a royal forest by William the Conqueror, fea ...
today. They were so prolific, culling had to be introduced in the 1930s to control their numbers.


Hunting

Across its original range and in many areas to which it has been introduced, the sika is regarded as a particularly prized and elusive sportsman's quarry. In Britain, Ireland, and mainland Europe, sika display very different survival strategies and escape tactics from the indigenous deer. They have a marked tendency to use concealment in circumstances when red deer, for example, would flee, and have been seen to squat and lie belly-flat when danger threatens. In the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isl ...
, sika are widely regarded as a serious threat to new and established woodlands, and public and private forestry bodies adopt policies of rigorous year-round culling.http://www.nonnativespecies.org/downloadDocument.cfm?id=355 The main predators of sika deer include
tiger The tiger (''Panthera tigris'') is the largest living Felidae, cat species and a member of the genus ''Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily pr ...
s,
wolves The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly un ...
,
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, ...
s, and
brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is k ...
s.
Lynx A lynx is a type of wild cat. Lynx may also refer to: Astronomy * Lynx (constellation) * Lynx (Chinese astronomy) * Lynx X-ray Observatory, a NASA-funded mission concept for a next-generation X-ray space observatory Places Canada * Lynx, ...
and
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird ...
s target fawns.


Velvet antler

Velvet antler (dried immature
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on ...
s) is a popular ingredient in
traditional Chinese medicine Traditional Chinese medicine (TCM) is an alternative medical practice drawn from traditional medicine in China. It has been described as "fraught with pseudoscience", with the majority of its treatments having no logical mechanism of acti ...
, and sika in China were domesticated long ago for the antler trade, along with several other species. In Taiwan, both Formosan sika deer and Formosan
sambar deer The sambar (''Rusa unicolor'') is a large deer native to the Indian subcontinent and Southeast Asia that is listed as a vulnerable species on the IUCN Red List since 2008. Populations have declined substantially due to severe hunting, local ins ...
(''Cervus unicolor swinhoei'') have been farmed for velvet antlers. Japan is the only country in eastern Asia where sika deer were not farmed for velvet antlers. Other deer raised for the antler trade were
Thorold's deer Thorold's deer (''Cervus albirostris'')Pitraa, Fickela, Meijaard, Groves (2004). ''Evolution and phylogeny of old world deer.'' Molecular Phylogenetics and Evolution 33: 880–895. is a threatened species of deer found in grassland, shrubland, an ...
(''Cervus albirostris''),
central Asian red deer The Central Asian red deer (''Cervus hanglu''), also known as the Tarim red deer is a deer species native to Central Asia, where it used to be widely distributed, but is scattered today with small population units in several countries. It has be ...
(''Cervus canadensis affinis''), and American elk (''Cervus canadensis canadensis'').


Cultural significance

In
Shinto Shinto () is a religion from Japan. Classified as an East Asian religion by scholars of religion, its practitioners often regard it as Japan's indigenous religion and as a nature religion. Scholars sometimes call its practitioners ''Shint ...
, the ''Shika'' Deer is considered a kind of messenger between mortals and the ''
kami are the deities, divinities, spirits, phenomena or "holy powers", that are venerated in the Shinto religion. They can be elements of the landscape, forces of nature, or beings and the qualities that these beings express; they can also be the sp ...
''.


See also

*
Deer of Great Britain Six species of deer are living wild in Great Britain: Scottish red deer, roe deer, fallow deer, sika deer, Reeves's muntjac, and Chinese water deer. Of those, Scottish red and roe deer are native and have lived in the isles throughout the Holocen ...
*
Nara Park is a public park located in the city of Nara, Japan, at the foot of Mount Wakakusa. Established in 1880, it is one of the oldest parks in Japan. Administratively, the park is under the control of Nara Prefecture. The park is one of the "Places o ...


References

* * Igota, H., Sakagura, M., Uno, H., Kaji, K., Maneko, M., Akamatsu, R., & Maekawa (in press). "Seasonal patterns of female sika deer in eastern Hokkaidō, Japan." ''Ecological Research'', 19.


Further reading

* O'Brien, D.J., Rooney, S.M. and Hayden, T.J. (September 2009). "A differential vulnerability to hunting between the sexes in Sika-type calves". ''I. Nat. J.'' 30: 7–9. .


External links


Nara's Sika Deer, from The Official Nara Travel Guide





Sika Deer in New Zealand and their distribution.
{{Authority control nippon Mammals of East Asia Mammals of Southeast Asia Mammals of China Mammals of Japan Mammals of Korea Mammals of Russia Mammals of Taiwan Mammals of Vietnam Mammals described in 1838 Taxa named by Coenraad Jacob Temminck Articles containing video clips