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The second voyage of HMS ''Beagle'', from 27 December 1831 to 2 October 1836, was the second survey expedition of HMS ''Beagle'', under captain Robert FitzRoy who had taken over command of the ship on its first voyage after the previous captain, Pringle Stokes, committed suicide. FitzRoy had thought of the advantages of having someone onboard who could investigate
geology Geology () is a branch of natural science concerned with Earth and other Astronomical object, astronomical objects, the features or rock (geology), rocks of which it is composed, and the processes by which they change over time. Modern geology ...
, and sought a naturalist to accompany them as a supernumerary. At the age of 22, the graduate
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
hoped to see the tropics before becoming a parson and accepted the opportunity. He was greatly influenced by reading Charles Lyell's '' Principles of Geology'' during the voyage. By the end of the expedition, Darwin had made his name as a geologist and
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
collector and the publication of his journal (later known as '' The Voyage of the Beagle'') gave him wide renown as a writer. ''Beagle'' sailed across the Atlantic Ocean, and then carried out detailed hydrographic surveys around the coasts of southern South America, returning via Tahiti and Australia after having
circumnavigated Circumnavigation is the complete navigation around an entire island, continent, or astronomical body (e.g. a planet or moon). This article focuses on the circumnavigation of Earth. The first recorded circumnavigation of the Earth was the ...
the Earth. The initial offer to Darwin told him the voyage would last two years; it lasted almost five. Darwin spent most of this time exploring on land: three years and three months land, 18 months at sea. Early in the voyage, Darwin decided that he could write a geology book, and he showed a gift for theorising. At Punta Alta in Argentina, he made a major find of gigantic fossils of extinct mammals, then known from very few specimens. He collected and made detailed observations of plants and animals. His findings undermined his belief in the doctrine that species are fixed, and provided the basis for ideas which came to him when back in England, leading to his theory of
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
by
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Cha ...
.


Aims of the expedition

When the
Napoleonic Wars The Napoleonic Wars (1803–1815) were a series of major global conflicts pitting the French Empire and its allies, led by Napoleon I, against a fluctuating array of European states formed into various coalitions. It produced a period of Fre ...
ended in 1815, the ''
Pax Britannica ''Pax Britannica'' (Latin for "British Peace", modelled after '' Pax Romana'') was the period of relative peace between the great powers during which the British Empire became the global hegemonic power and adopted the role of a "global poli ...
'' saw seafaring nations competing in colonisation and rapid industrialisation. The logistics of supply and growing commerce needed reliable information about sea routes, but existing
nautical chart A nautical chart is a graphic representation of a sea area and adjacent coastal regions. Depending on the scale of the chart, it may show depths of water and heights of land ( topographic map), natural features of the seabed, details of the co ...
s were incomplete and inaccurate.
Spanish American wars of independence The Spanish American wars of independence (25 September 1808 – 29 September 1833; es, Guerras de independencia hispanoamericanas) were numerous wars in Spanish America with the aim of political independence from Spanish rule during the early ...
ended Spain's monopoly over trade, and the UK's 1825 commercial treaty with Argentina recognised its independence, increasing the naval and commercial significance of the east coast of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
. The
Admiralty Admiralty most often refers to: *Admiralty, Hong Kong *Admiralty (United Kingdom), military department in command of the Royal Navy from 1707 to 1964 *The rank of admiral *Admiralty law Admiralty can also refer to: Buildings * Admiralty, Traf ...
instructed Commander King to make an accurate hydrographic survey of "the Southern Coasts of the Peninsula of South America, from the southern entrance of the River Plata, round to Chilóe; and of Tierra del Fuego". As Darwin wrote of his voyage, "The object of the expedition was to complete the survey of Patagonia and Tierra del Fuego, commenced under Captain King in 1826 to 1830—to survey the shores of Chile, Peru, and of some islands in the Pacific—and to carry a chain of chronometrical measurements round the World." The expeditions also had diplomatic objectives, visiting disputed territories. An Admiralty memorandum set out the detailed instructions. The first requirement was to resolve disagreements in the earlier surveys about the
longitude Longitude (, ) is a geographic coordinate that specifies the east– west position of a point on the surface of the Earth, or another celestial body. It is an angular measurement, usually expressed in degrees and denoted by the Greek let ...
of
Rio de Janeiro Rio de Janeiro ( , , ; literally 'River of January'), or simply Rio, is the capital of the state of the same name, Brazil's third-most populous state, and the second-most populous city in Brazil, after São Paulo. Listed by the GaWC as a b ...
, which was essential as the base point for meridian distances. The accurate
marine chronometer A marine chronometer is a precision timepiece that is carried on a ship and employed in the determination of the ship's position by celestial navigation. It is used to determine longitude by comparing Greenwich Mean Time (GMT), or in the mode ...
s needed to determine longitude had only become affordable since 1800; ''Beagle'' carried 22 chronometers to allow corrections. The ship was to stop at specified points for a four-day rating of the chronometers and to check them by astronomical observations: it was essential to take observations at Porto Praya and Fernando de Noronha to calibrate against the previous surveys of
William Fitzwilliam Owen Vice Admiral William Fitzwilliam Owen (17 September 1774 – 3 November 1857), was a British naval officer and explorer. He is best known for his exploration of the west and east African coasts, discovery of the Seaflower Channel off the co ...
and Henry Foster. It was important to survey the extent of the
Abrolhos Archipelago The Abrolhos Archipelago () are a group of 5 small islands with coral reefs off the southern coast of Bahia state in the northeast of Brazil, between 17º25’—18º09’ S and 38º33’—39º05’ W. Caravelas is the nearest town. Their name ...
reefs, shown incorrectly in Albin Roussin's survey, then proceed to Rio de Janeiro to decide the exact longitude of
Villegagnon Island Villegagnon Island (former Serigipe Island—original Portuguese: ''Ilha de Villegagnon''—also known in English as: Villegaignon Island, Island of Villegagnon or Island of Villegaignon) is located near the mouth of the large Guanabara Bay, in th ...
. The real work of the survey was then to commence south of the Río de la Plata, with return trips to
Montevideo Montevideo () is the capital and largest city of Uruguay. According to the 2011 census, the city proper has a population of 1,319,108 (about one-third of the country's total population) in an area of . Montevideo is situated on the southern co ...
for supplies; details were given of priorities, including surveying Tierra del Fuego and approaches to harbours on the
Falkland Islands The Falkland Islands (; es, Islas Malvinas, link=no ) is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and about from Cape Dubouze ...
. The west coast was then to be surveyed as far north as time and resources permitted. The commander would then determine his own route west: season permitting, he could survey the
Galápagos Islands The Galápagos Islands ( Spanish: , , ) are an archipelago of volcanic islands. They are distributed on each side of the equator in the Pacific Ocean, surrounding the centre of the Western Hemisphere, and are part of the Republic of Ecuad ...
. Then, ''Beagle'' was to proceed to Point Venus, Tahiti, and on to
Port Jackson Port Jackson, consisting of the waters of Sydney Harbour, Middle Harbour, North Harbour and the Lane Cove and Parramatta Rivers, is the ria or natural harbour of Sydney, New South Wales, Australia. The harbour is an inlet of the Tasman S ...
, Australia, which were known points to verify the chronometers. No time was to be wasted on elaborate drawings; charts and plans should have notes and simple views of the land as seen from the sea showing measured heights of hills. Continued records of tides and
meteorological Meteorology is a branch of the atmospheric sciences (which include atmospheric chemistry and physics) with a major focus on weather forecasting. The study of meteorology dates back millennia, though significant progress in meteorology did not ...
conditions were also required. An additional suggestion was for a geological survey of a circular coral
atoll An atoll () is a ring-shaped island, including a coral rim that encircles a lagoon partially or completely. There may be coral islands or cays on the rim. Atolls are located in warm tropical or subtropical oceans and seas where corals can gro ...
in the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the conti ...
including its profile and of tidal flows, to investigate the formation of such coral reefs.


Context and preparations

The previous survey expedition to South America involved HMS ''Adventure'' and HMS ''Beagle'' under the overall command of the Australian Commander
Phillip Parker King Rear Admiral Phillip Parker King, FRS, RN (13 December 1791 – 26 February 1856) was an early explorer of the Australian and Patagonian coasts. Early life and education King was born on Norfolk Island, to Philip Gidley King and Anna ...
. During the survey, ''Beagle'' captain, Pringle Stokes, committed suicide and command of the ship was given to the young aristocrat Robert FitzRoy, a nephew of George FitzRoy, 4th Duke of Grafton. When a ship's boat was taken by the natives of Tierra del Fuego, FitzRoy tried taking some of them hostage, and after this failed he got occupants of a canoe to put another on the ship in exchange for buttons. He brought four of them back to England to be given a Christian education, with the idea that they could eventually become missionaries. One died of
smallpox Smallpox was an infectious disease caused by variola virus (often called smallpox virus) which belongs to the genus Orthopoxvirus. The last naturally occurring case was diagnosed in October 1977, and the World Health Organization (WHO) c ...
. After ''Beagle'' return to Devonport dockyard on 14 October 1830, Captain King retired. The 27-year-old FitzRoy had hopes of commanding a second expedition to continue the South American survey, but when he heard that the Lords of the Admiralty no longer supported this, he grew concerned about how to return the Fuegians. He made an agreement with the owner of a small merchant-vessel to take himself and five others back to South America, but a kind uncle heard of this and contacted the Admiralty. Soon afterwards, FitzRoy heard that he was to be appointed commander of HMS ''Chanticleer'' to go to Tierra del Fuego, but due to her poor condition, ''Beagle'' was substituted. On 27 June 1831, FitzRoy was commissioned as commander of the voyage, and Lieutenants John Clements Wickham and
Bartholomew James Sulivan Admiral Sir Bartholomew James Sulivan, (18 November 1810 – 1 January 1890) was a British naval officer and hydrographer. He was a leading advocate of the value of nautical surveying in relation to naval operations. Sulivan was born at Mylor, ...
were both appointed. Captain
Francis Beaufort Rear-Admiral Sir Francis Beaufort (; 27 May 1774 – 17 December 1857) was an Irish hydrographer, rear admiral of the Royal Navy, and creator of the Beaufort cipher and the Beaufort scale. Early life Francis Beaufort was descende ...
, the Hydrographer of the Admiralty, was invited to decide on the use that could be made of the voyage to continue the survey, and he discussed with FitzRoy plans for a voyage of several years, including a continuation of the trip around the world to establish median distances. ''Beagle'' was commissioned on 4 July 1831, under the command of Captain FitzRoy, who promptly spared no expense in having ''Beagle'' extensively refitted. ''Beagle'' was immediately taken into dock for extensive rebuilding and refitting. As she required a new deck, FitzRoy had the upper-deck raised considerably, by aft and forward. The ''Cherokee''-class brig-sloops had the reputation of being "coffin brigs", which handled badly and were prone to sinking. By helping the decks to drain more quickly with less water collecting in the gunnels, the raised deck gave ''Beagle'' better handling and made her less liable to become top-heavy and capsize. Additional sheathing to the hull added about seven tons to her
burthen Burden or burthen may refer to: People * Burden (surname), people with the surname Burden Places * Burden, Kansas, United States * Burden, Luxembourg Arts, entertainment, and media Films * ''Burden'' (2018 film), an American drama film * '' ...
and perhaps fifteen to her displacement. The ship was one of the first to test the lightning conductor invented by William Snow Harris. FitzRoy obtained five examples of the '' Sympiesometer'', a kind of mercury-free barometer patented by
Alexander Adie Alexander James Adie FRSE MWS (1775, Edinburgh – 4 December 1858, Edinburgh) was a Scottish maker of medical instruments, optician and meteorologist. He was the inventor of the sympiesometer, patented in 1818. Life He was born the son of Jo ...
and favoured by FitzRoy as giving the accurate readings required by the Admiralty. In addition to its officers and crew, ''Beagle'' carried several supernumeraries, passengers without an official position. FitzRoy employed a mathematical instrument maker to maintain his 22 marine chronometers kept in his cabin, as well as engaging the artist/draughtsman
Augustus Earle Augustus Earle (1793–1838) was a British painter. Unlike earlier artists who worked outside Europe and were employed on voyages of exploration or worked abroad for wealthy, often aristocratic patrons, Earle was able to operate quite indepe ...
to go in a private capacity. The three Fuegians taken on the previous voyage were going to be returned to Tierra del Fuego on ''Beagle'' together with the missionary Richard Matthews.


Naturalist and geologist

For
Beaufort Beaufort may refer to: People and titles * Beaufort (surname) * House of Beaufort, English nobility * Duke of Beaufort (England), a title in the peerage of England * Duke of Beaufort (France), a title in the French nobility Places Polar regions * ...
and the leading
Cambridge Cambridge ( ) is a university city and the county town in Cambridgeshire, England. It is located on the River Cam approximately north of London. As of the 2021 United Kingdom census, the population of Cambridge was 145,700. Cambridge bec ...
"gentlemen of science" the opportunity for a naturalist to join the expedition fitted with their drive to revitalise British government policy on science. This elite disdained research done for money and felt that
natural philosophy Natural philosophy or philosophy of nature (from Latin ''philosophia naturalis'') is the philosophical study of physics, that is, nature and the physical universe. It was dominant before the development of modern science. From the ancien ...
was for gentlemen, not
tradesmen A tradesman, tradeswoman, or tradesperson is a skilled worker that specializes in a particular trade (occupation or field of work). Tradesmen usually have work experience, on-the-job training, and often formal vocational education in contrast ...
. The officer class of the Army and Navy provided a way to ascend this hierarchy; the ship's surgeon often collected specimens on voyages, and Robert McCormick had secured the position on ''Beagle'' after taking part in earlier expeditions and studying natural history. A sizeable collection had considerable social value, attracting wide public interest, and McCormick aspired to fame as an exploring naturalist. Collections made by the ship's surgeon and other officers were government property, though the Admiralty was not consistent on this, and went to important London establishments, usually the
British Museum The British Museum is a public museum dedicated to human history, art and culture located in the Bloomsbury area of London. Its permanent collection of eight million works is among the largest and most comprehensive in existence. It docum ...
. The Admiralty instructions for the first voyage had required officers "to use their best diligence in increasing the Collections in each ship: the whole of which must be understood to belong to the Public", but on the second voyage this requirement was omitted, and the officers were free to keep all the specimens for themselves. FitzRoy's journal written during the first voyage noted that, while investigating magnetic rocks near the Barbara Channel, he regretted "that no person in the vessel was skilled in
mineralogy Mineralogy is a subject of geology specializing in the scientific study of the chemistry, crystal structure, and physical (including optical) properties of minerals and mineralized artifacts. Specific studies within mineralogy include the proce ...
, or at all acquainted with
geology Geology () is a branch of natural science concerned with Earth and other Astronomical object, astronomical objects, the features or rock (geology), rocks of which it is composed, and the processes by which they change over time. Modern geology ...
", to make use of the opportunity of "ascertaining the nature of the rocks and earths" of the areas surveyed. FitzRoy decided that on any similar future expedition, he would "endeavour to carry out a person qualified to examine the land; while the officers, and myself, would attend to
hydrography Hydrography is the branch of applied sciences which deals with the measurement and description of the physical features of oceans, seas, coastal areas, lakes and rivers, as well as with the prediction of their change over time, for the prima ...
." This indicated a need for a naturalist qualified to examine geology, who would spend considerable periods onshore away from the ship. McCormick lacked expertise in geology and had to attend to his duties on the ship. FitzRoy knew that commanding a ship could involve stress and loneliness. He was aware of his uncle
Viscount Castlereagh A viscount ( , for male) or viscountess (, for female) is a title used in certain European countries for a noble of varying status. In many countries a viscount, and its historical equivalents, was a non-hereditary, administrative or judici ...
's suicide due to stress from overwork, as well as Captain Stokes's suicide. This was to be the first time that FitzRoy would be fully in charge of a ship with no commanding officer or second captain to consult. It has been suggested that he felt the need for a gentleman companion who shared his scientific interests and could dine with him as an equal, although there is no direct evidence to support this. Professor John Stevens Henslow described the position "more as a companion than a mere collector", but this was an assurance that FitzRoy would treat his guest as a gentleman naturalist. Several other ships at this period carried unpaid civilians as naturalists. Early in August, FitzRoy discussed this position with Beaufort, who had a scientific network of friends at the
University of Cambridge , mottoeng = Literal: From here, light and sacred draughts. Non literal: From this place, we gain enlightenment and precious knowledge. , established = , other_name = The Chancellor, Masters and Schola ...
. At Beaufort's request, mathematics lecturer George Peacock wrote from London to Henslow about this "rare opportunity for a naturalist", saying that an "offer has been made to me to recommend a proper person to go out as a naturalist with this expedition", and suggesting the Reverend Leonard Jenyns. Though Jenyns nearly accepted and even packed his clothes, he had concerns about his obligations as vicar of
Swaffham Bulbeck Swaffham Bulbeck is a small village in East Cambridgeshire, England. Swaffham Bulbeck is located about from the city of Cambridge, and from the famous racing town of Newmarket. The parish of Swaffham Bulbeck is part of the Diocese of Ely an ...
and about his health, therefore Jenyns declined the offer. Henslow briefly thought of going, but his wife "looked so miserable" that he quickly dropped the idea. Both recommended bringing the 22-year-old
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
, who was on a geology field trip with
Adam Sedgwick Adam Sedgwick (; 22 March 1785 – 27 January 1873) was a British geologist and Anglican priest, one of the founders of modern geology. He proposed the Cambrian and Devonian period of the geological timescale. Based on work which he did on ...
. He had just completed the ordinary
Bachelor of Arts Bachelor of arts (BA or AB; from the Latin ', ', or ') is a bachelor's degree awarded for an undergraduate program in the arts, or, in some cases, other disciplines. A Bachelor of Arts degree course is generally completed in three or four yea ...
degree which was a prerequisite for his intended career as a parson.


Offer of place to Darwin

Darwin fitted well the expectations of a gentleman natural philosopher and was well trained as a naturalist. When he had studied geology in his second year at Edinburgh, he had found it dull, but from Easter to August 1831, he learned a great deal with Sedgwick and developed a strong interest during their geological field trip. On 24 August Henslow wrote to Darwin:
...that I consider you to be the best qualified person I know of who is likely to undertake such a situation— I state this not on the supposition of yr. being a finished Naturalist, but as amply qualified for collecting, observing, & noting any thing worthy to be noted in Natural History. Peacock has the appointment at his disposal & if he can not find a man willing to take the office, the opportunity will probably be lost— Capt. F. wants a man (I understand) more as a companion than a mere collector & would not take any one however good a Naturalist who was not recommended to him likewise as a ''gentleman''. ... The Voyage is to last 2 yrs. & if you take plenty of Books with you, any thing you please may be done ... there never was a finer chance for a man of zeal & spirit... Don't put on any modest doubts or fears about your disqualifications for I assure you I think you are the very man they are in search of.
The letter went first to George Peacock, who quickly forwarded it to Darwin with further details, confirming that the "ship sails about the end of September". Peacock had discussed the offer with
Beaufort Beaufort may refer to: People and titles * Beaufort (surname) * House of Beaufort, English nobility * Duke of Beaufort (England), a title in the peerage of England * Duke of Beaufort (France), a title in the French nobility Places Polar regions * ...
, "he entirely approves of it & you may consider the situation as at your absolute disposal". When Darwin returned home from the field trip late on 29 August and opened the letters, his father objected strongly to the voyage so, the next day, he wrote declining the offer and left to go shooting at the estate of his uncle
Josiah Wedgwood II Josiah Wedgwood II (3 April 1769 – 12 July 1843), the son of the English potter Josiah Wedgwood, continued his father's firm and was a Member of Parliament (MP) for Stoke-upon-Trent from 1832 to 1835. He was an abolitionist, and detested slav ...
. With Wedgwood's help, Darwin's father was persuaded to relent and fund his son's expedition, and on Thursday 1 September, Darwin wrote to Beaufort accepting the offer. That day, Beaufort wrote to tell FitzRoy that his friend Peacock had "succeeded in getting a '
Savant Savant syndrome () is a rare condition in which someone with significant mental disabilities demonstrates certain abilities far in excess of average. The skills that savants excel at are generally related to memory. This may include rapid calc ...
' for you—A Mr Darwin grandson of the well known philosopher and poet—full of zeal and enterprize and having contemplated a voyage on his own account to S. America". On Friday, Darwin left for Cambridge, where he, the next day, got advice on preparations of the voyage and references to experts by Henslow. Alexander Charles Wood (an undergraduate whose tutor was Peacock) wrote from Cambridge to his cousin FitzRoy to recommend Darwin. Around midday on Sunday 4 September, Wood received FitzRoy's response, "straightforward and gentlemanlike" but strongly against Darwin joining the expedition; both Darwin and Henslow then "gave up the scheme". Darwin went to London anyway, and next morning met FitzRoy, who explained that he had promised the place to his friend Mr. Chester (possibly the novelist Harry Chester), but Chester had turned it down in a letter received not five minutes before Darwin arrived. FitzRoy emphasised the difficulties, including cramped conditions and plain food. Darwin would be on the Admiralty's books to get provisions (worth £40 a year) and, like the ship's officers and captain, would pay £30 a year towards the
mess The mess (also called a mess deck aboard ships) is a designated area where military personnel socialize, eat and (in some cases) live. The term is also used to indicate the groups of military personnel who belong to separate messes, such as the o ...
bill. Including outfitting, the cost to him was unlikely to reach £500. The ship would sail on 10 October, and would probably be away for three years. They talked and dined together, and soon found each other agreeable. The Tory FitzRoy had been cautious at the prospect of companionship with this unknown young gentleman of Whig background, and later admitted that his letter to Wood was "to throw cold water on the scheme" in "a sudden horror of the chances of having somebody he should not like on board". He half-seriously told Darwin later that, as "an ardent disciple of Lavater", he had nearly rejected Darwin on the
phrenological Phrenology () is a pseudoscience which involves the measurement of bumps on the skull to predict mental traits.Wihe, J. V. (2002). "Science and Pseudoscience: A Primer in Critical Thinking." In ''Encyclopedia of Pseudoscience'', pp. 195–203. C ...
basis that the shape (or physiognomy) of Darwin's nose indicated a lack of determination.


Darwin's preparations

While he continued to get acquainted with FitzRoy, going shopping together, Darwin rushed around to arrange his supplies and equipment. He took advice from experts on specimen preservation including William Yarrell at the
Zoological Society of London The Zoological Society of London (ZSL) is a charity devoted to the worldwide conservation of animals and their habitats. It was founded in 1826. Since 1828, it has maintained the London Zoo, and since 1931 Whipsnade Park. History On 29 ...
, Robert Brown at the British Museum, Captain
Phillip Parker King Rear Admiral Phillip Parker King, FRS, RN (13 December 1791 – 26 February 1856) was an early explorer of the Australian and Patagonian coasts. Early life and education King was born on Norfolk Island, to Philip Gidley King and Anna ...
who led the first expedition, and invertebrate anatomist Robert Edmond Grant who had tutored Darwin at Edinburgh. Yarrell gave invaluable advice and bargained with shopkeepers, so Darwin paid £50 for two pistols and a rifle, while FitzRoy had spent £400 on firearms. On Sunday, 11 September, FitzRoy and Darwin took the
steam packet Generally, packet trade is any regularly scheduled cargo, passenger and mail trade conducted by ship. The ships are called "packet boats" as their original function was to carry mail. A "packet ship" was originally a vessel employed to carry post ...
for Portsmouth. Darwin was not seasick and had a pleasant "sail of three days". For the first time, he saw the "very small" cramped ship, met the officers, and was glad to get a large cabin, shared with the assistant surveyor John Lort Stokes. On Friday, Darwin rushed back to London, "250 miles in 24 hours", and on via Cambridge and St. Albans, travelling on the Wonder coach all day on 22 September to arrive in Shrewsbury that evening, then after a last brief visit to family and friends left for London on 2 October. Delays to ''Beagle'' gave Darwin an extra week to consult experts and complete packing his baggage. After sending his heavy goods down by steam packet, he took the coach along with
Augustus Earle Augustus Earle (1793–1838) was a British painter. Unlike earlier artists who worked outside Europe and were employed on voyages of exploration or worked abroad for wealthy, often aristocratic patrons, Earle was able to operate quite indepe ...
and arrived at Devonport on 24 October. The geologist Charles Lyell asked FitzRoy to record observations on geological features such as erratic boulders. Before they left England, FitzRoy gave Darwin a copy of the first volume of Lyell's '' Principles of Geology'' which explained features as the outcome of a gradual process taking place over extremely long periods of time. In his autobiography, Darwin recalled Henslow giving advice at this time to obtain and study the book, "but on no account to accept the views therein advocated". Darwin's position as a naturalist on board was as a self-funded guest with no official appointment, and he could leave the voyage at any suitable stage. At the outset, George Peacock had advised that "The Admiralty are not disposed to give a salary, though they will furnish you with an official appointment & every : if a salary should be required however I am inclined to think that it would be granted". Far from wanting this, Darwin's concern was to maintain control over his collection. He was even reluctant to be on the Admiralty's books for
victuals Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is ...
until he got assurances from FitzRoy and Beaufort that this would not affect his rights to assign his specimens. Beaufort initially thought specimens ought to go to the British Museum, but Darwin had heard of many left waiting to be described, including botanical specimens from the first ''Beagle'' voyage. Beaufort assured him that he "should have no difficulty" as long as he "presented them to some public body" such as the Zoological or Geological societies. Henslow had set up the small
Cambridge Philosophical Society The Cambridge Philosophical Society (CPS) is a scientific society at the University of Cambridge. It was founded in 1819. The name derives from the medieval use of the word philosophy to denote any research undertaken outside the fields of l ...
museum, Darwin told him that new finds should go to the "largest & most central collection" rather than a "Country collection, let it be ever so good", but soon expressed "hope to be able to assist the Philosoph. Society" with some specimens. FitzRoy arranged transport of specimens to England as official cargo on the
Admiralty Admiralty most often refers to: *Admiralty, Hong Kong *Admiralty (United Kingdom), military department in command of the Royal Navy from 1707 to 1964 *The rank of admiral *Admiralty law Admiralty can also refer to: Buildings * Admiralty, Traf ...
Packet Service, at no cost to Darwin even though it was his private collection. Henslow agreed to store them at Cambridge, and Darwin confirmed with him arrangements for land carriage from the port, to be funded by Darwin's father.


Darwin's work on the expedition

The captain had to record his survey in painstaking paperwork, and Darwin too kept a daily log as well as detailed notebooks of his finds and speculations, and a diary which became his journal. Darwin's notebooks show complete professionalism that he had probably learnt at the
University of Edinburgh The University of Edinburgh ( sco, University o Edinburgh, gd, Oilthigh Dhùn Èideann; abbreviated as ''Edin.'' in post-nominals) is a public research university based in Edinburgh, Scotland. Granted a royal charter by King James VI in 1 ...
when making natural history notes while exploring the shores of the Firth of Forth with his brother Erasmus in 1826 and studying marine invertebrates with Robert Edmund Grant for a few months in 1827. Darwin had also collected
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s at Cambridge, but he was a novice in all other areas of natural history. During the voyage, Darwin investigated small invertebrates while collecting specimens of other creatures for experts to examine and describe once ''Beagle'' had returned to England. More than half of his carefully organised zoology notes deal with marine invertebrates. The notes also record closely reasoned interpretations of what he found about their complex internal anatomy while dissecting specimens under his microscope and small experiments on their response to stimulation. His onshore observations included intense, analytical comments on possible reasons for the behaviour, distribution, and relation to their environment of the creatures he saw. He made good use of the ship's excellent library of books on natural history but continually questioned their correctness. Geology was Darwin's "principal pursuit" on the expedition, and his notes on that subject were almost four times larger than his zoology notes, although he kept extensive records on both. During the voyage, he wrote to his sister that "there is nothing like geology; the pleasure of the first days partridge shooting or first days hunting cannot be compared to finding a fine group of fossil bones, which tell their story of former times with almost a living tongue". To him, investigating geology brought reasoning into play and gave him opportunities for theorising.


Voyage

Charles Darwin had been told that ''Beagle'' was expected to sail about the end of September 1831, but fitting out took longer. The Admiralty Instructions were received on 14 November, and on 23 November, she was moved to anchorage, ready to depart. Repeated Westerly gales caused delays, and forced them to turn back after departing on 10 and 21 December. Drunkenness at Christmas lost another day. Finally, on the morning of 27 December, ''Beagle'' left its anchorage in the Barn Pool, under Mount Edgecumbe on the west side of Plymouth Sound and set out on its surveying expedition.


Atlantic islands

''Beagle'' touched at
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
for a confirmed position without stopping. Then on 6 January, it reached
Tenerife Tenerife (; ; formerly spelled ''Teneriffe'') is the largest and most populous island of the Canary Islands. It is home to 43% of the total population of the Archipelago, archipelago. With a land area of and a population of 978,100 inhabitant ...
in the
Canary Islands The Canary Islands (; es, :es:Canarias, Canarias, ), also known informally as the Canaries, are a Spanish Autonomous communities of Spain, autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to ...
but was quarantined there because of
cholera Cholera is an infection of the small intestine by some strains of the bacterium '' Vibrio cholerae''. Symptoms may range from none, to mild, to severe. The classic symptom is large amounts of watery diarrhea that lasts a few days. Vomiting an ...
in England. Although tantalisingly near to the town of Santa Cruz, to Darwin's intense disappointment, they were denied landing. With improving weather conditions, they sailed on. On 10 January, Darwin tried out a plankton net he had devised to be towed behind the ship—only the second recorded use of such a net (after use by John Vaughan Thompson in 1816). Next day, he noted the great number of animals collected far from land and wrote: "Many of these creatures so low in the scale of nature are most exquisite in their forms & rich colours. — It creates a feeling of wonder that so much beauty should be apparently created for such little purpose." Six days later, they made their first landing at Praia on the volcanic island of
Santiago Santiago (, ; ), also known as Santiago de Chile, is the capital and largest city of Chile as well as one of the largest cities in the Americas. It is the center of Chile's most densely populated region, the Santiago Metropolitan Region, whos ...
in the
Cape Verde , national_anthem = () , official_languages = Portuguese , national_languages = Cape Verdean Creole , capital = Praia , coordinates = , largest_city = capital , demonym ...
Islands. It is here that Darwin's description in his published '' Journal'' begins. His initial impression was of a desolate and sterile volcanic island. However, upon visiting the town, he came to a deep valley where he "first saw the glory of tropical vegetation" and had "a glorious day", finding overwhelming novelty in the sights and sounds. FitzRoy set up tents and an observatory on Quail Island to determine the exact position of the islands, while Darwin collected numerous sea animals, delighting in vivid tropical corals in tidal pools, and investigating the geology of Quail Island. Though Daubeny's book in ''Beagle'' library described the volcanic geology of the Canary Islands, it said that the structure of the Cape Verde Islands was "too imperfectly known". Darwin saw Quail Island as his key to understanding the structure of St. Jago and made careful studies of its
stratigraphy Stratigraphy is a branch of geology concerned with the study of rock layers ( strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithost ...
in the way he had learnt from Adam Sedgwick. He collected specimens and described a white layer of hard white rock formed from crushed coral and seashells lying between layers of black volcanic rocks, and noted a similar white layer running horizontally in the black cliffs of St. Jago at above sea level. The seashells were, as far as he could tell, "the same as those of present day". He speculated that in geologically recent times a lava flow had covered this shell sand on the sea bed, and then the strata had slowly risen to their present level. Charles Lyell's ''Principles of Geology'' presented a thesis of gradual rising and falling of the Earth's crust illustrated by the changing levels of the Temple of Serapis. Darwin implicitly supported Lyell by remarking that "Dr. Daubeny when mentioning the present state of the temple of Serapis. doubts the possibility of a surface of country being raised without cracking buildings on it. – I feel sure at St Jago in some places a town might have been raised without injuring a house." Later, in his first letter to Henslow, he wrote that "The geology was preeminently interesting & I believe quite new: there are some facts on a large scale of upraised coast ... that would interest Mr. Lyell." While still on the island, Darwin was inspired to think of writing a book on geology, and later wrote of "seeing a thing never seen by Lyell, one yet saw it partially through his eyes". ''Beagle'' surgeon Robert McCormick sought fame and fortune as an explorer. When they first met at the start of the voyage, Darwin had commented that "My friend cCormickis an ass, but we jog on very amicably". They walked into the countryside of St. Jago together, and Darwin, influenced by Lyell, found the surgeon's approach old-fashioned. They found a remarkable
baobab ''Adansonia'' is a genus made up of eight species of medium-to-large deciduous trees known as baobabs ( or ). They are placed in the Malvaceae family, subfamily Bombacoideae. They are native to Madagascar, mainland Africa, and Australia.T ...
tree, which FitzRoy measured and sketched. Darwin went on subsequent "riding expeditions" with
Benjamin Bynoe Benjamin Bynoe (1803–1865) was surgeon on the voyages of HMS ''Beagle'' who made collections of plants and animals at the western and northern coasts of Australia. Born in Barbados in 1803, Benjamin Bynoe was accepted by the Royal College of S ...
and Rowlett to visit
Ribeira Grande Ribeira Grande may refer to the following places: Cape Verde *Ribeira Grande (stream), a stream on the island of Santo Antão *Ribeira Grande, Cape Verde, a town on the island of Santo Antão *Ribeira Grande, Cape Verde (municipality), a municipali ...
and St Domingo. FitzRoy extended their stay to 23 days to complete his measurements of
magnetism Magnetism is the class of physical attributes that are mediated by a magnetic field, which refers to the capacity to induce attractive and repulsive phenomena in other entities. Electric currents and the magnetic moments of elementary particles ...
. Darwin subsequently wrote to Henslow that his collecting included "several specimens of an Octopus, which possessed a most marvellous power of changing its colours; equalling any chamaelion, & evidently accommodating the changes to the colour of the ground which it passed over.—yellowish green, dark brown & red were the prevailing colours: this fact appears to be new, as far as I can find out." Henslow replied that "The fact is not new, but any fresh observations will be highly important." McCormick increasingly resented the favours FitzRoy gave to assist Darwin with collecting. On 16 February, FitzRoy landed a small party including himself and Darwin on St. Paul's Rocks, finding the seabirds so tame that they could be killed easily, while an exasperated McCormick was left circling the islets in a second small boat. That evening, novices were greeted by a pseudo- Neptune, and in the morning, they crossed the
equator The equator is a circle of latitude, about in circumference, that divides Earth into the Northern and Southern hemispheres. It is an imaginary line located at 0 degrees latitude, halfway between the North and South poles. The term can also ...
with the traditional line-crossing ceremony. Darwin had a special position as a guest and social equal of the captain, so junior officers called him "sir" until the captain dubbed Darwin ''Philos'' for "ship's philosopher", which became his suitably respectful nickname.


Surveying South America

In South America, ''Beagle'' carried out its survey work going to and fro along the coasts to allow careful measurement and rechecking. Darwin made long journeys inland with travelling companions from the locality. He spent much of the time away from the ship, returning by prearrangement when ''Beagle'' returned to ports where mail and newspapers were received, and Darwin's notes, journals, and collections sent back to England, via the Admiralty Packet Service. He had ensured that his collections were his own and, as prearranged, batches of his specimens were shipped to England, then taken by land carriage to Henslow in
Cambridge Cambridge ( ) is a university city and the county town in Cambridgeshire, England. It is located on the River Cam approximately north of London. As of the 2021 United Kingdom census, the population of Cambridge was 145,700. Cambridge bec ...
to await his return. The first batch was sent in August 1832, journey time varied considerably but all batches were eventually delivered. Several others on board, including FitzRoy and other officers, were able amateur naturalists, and they gave Darwin generous assistance as well as making collections for the Crown, which the Admiralty placed in the British Museum.


Tropical paradise and slavery

Due to heavy surf, they only stayed at Fernando de Noronha for a day to make the required observations, then FitzRoy pressed on to Bahia de Todos Santos, Brazil, to rate the chronometers and take on water. They reached the continent and arrived at the port on 28 February. Darwin was thrilled at the magnificent sight of "the town of
Bahia Bahia ( , , ; meaning "bay") is one of the 26 states of Brazil, located in the Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo, Minas Gerais, and Rio de Janeiro) and the 5th-larges ...
or St Salvador", with large ships at harbour scattered across the bay. On the next day, he was in "transports of pleasure" walking by himself in the tropical forest, and in "long naturalizing walks" with others continued to "add raptures to the former raptures". He found the sights of slavery offensive, and when FitzRoy defended the practice by describing a visit to a slaveowner whose slaves replied "no" on being asked by their master if they wished to be freed, Darwin suggested that answers in such circumstances were worthless. Enraged that his word had been questioned, FitzRoy lost his temper and banned Darwin from his company. The officers had nicknamed such outbursts "hot coffee", and within hours FitzRoy apologised, and asked Darwin to remain. Later, FitzRoy had to remain silent when Captain Paget visited them and recounted "facts about slavery so revolting" that refuted his claim. Surveying of sandbanks around the harbour was completed on 18 March, and the ship made its way down the coast to survey the extent and depths of the Abrolhos reefs, completing and correcting Roussin's survey. They manoeuvred ''Beagle'' into
Rio de Janeiro Rio de Janeiro ( , , ; literally 'River of January'), or simply Rio, is the capital of the state of the same name, Brazil's third-most populous state, and the second-most populous city in Brazil, after São Paulo. Listed by the GaWC as a b ...
harbour "in first rate style" on 4 April, with Darwin enthusiastically helping. Amidst excitement at opening letters from home, he was taken aback by news that his close friend Fanny Owen was engaged to marry
Biddulph Biddulph is a town in Staffordshire, England, north of Stoke-on-Trent and south-east of Congleton, Cheshire. Origin of the name Biddulph's name may come from Anglo-Saxon/Old English ''bī dylfe'' = "beside the pit or quarry". It may also ...
of
Chirk Castle Chirk Castle ( cy, Castell y Waun) is a Grade I listed castle located in Chirk, Wrexham County Borough, Wales. History The castle was built in 1295 by Roger Mortimer de Chirk, uncle of Roger Mortimer, 1st Earl of March as part of King Ed ...
.
Augustus Earle Augustus Earle (1793–1838) was a British painter. Unlike earlier artists who worked outside Europe and were employed on voyages of exploration or worked abroad for wealthy, often aristocratic patrons, Earle was able to operate quite indepe ...
showed Darwin round the town, and they found a delightful cottage for lodgings at
Botafogo Botafogo (local/standard alternative Brazilian Portuguese pronunciation: ) is a beachfront neighborhood (''bairro'') in Rio de Janeiro, Brazil. It is a mostly upper middle class and small commerce community, and is located between the hills o ...
. Darwin made arrangements with local estate owners, and on 8 April set off with them on a strenuous "riding excursion" to Rio Macaè. McCormick had made himself disagreeable to FitzRoy and
first lieutenant First lieutenant is a commissioned officer military rank in many armed forces; in some forces, it is an appointment. The rank of lieutenant has different meanings in different military formations, but in most forces it is sub-divided into a ...
Wickham, so was "invalided home", as he also was on other voyages. In his 1884 memoirs, he claimed he had been "very much disappointed in my expectations of carrying out my natural history pursuits, every obstacle having been placed in the way of my getting on shore and making collections". Assistant Surgeon
Benjamin Bynoe Benjamin Bynoe (1803–1865) was surgeon on the voyages of HMS ''Beagle'' who made collections of plants and animals at the western and northern coasts of Australia. Born in Barbados in 1803, Benjamin Bynoe was accepted by the Royal College of S ...
was made acting surgeon in his place. The required observations from
Villegagnon Island Villegagnon Island (former Serigipe Island—original Portuguese: ''Ilha de Villegagnon''—also known in English as: Villegaignon Island, Island of Villegagnon or Island of Villegaignon) is located near the mouth of the large Guanabara Bay, in th ...
at Rio showed a discrepancy of of longitude in the meridian distance from Bahia to Rio, compared to Roussin's results, and FitzRoy wrote telling Beaufort he would go back to check. On 24 April Darwin got back to the ship, next day his books, papers, and equipment suffered minor damage when the boat taking him to Botafogo cottage was swamped. He sent his sister his " commonplace Journal" to date, inviting criticisms, and decided to stay in the cottage with Earle while the ship went to Bahia. Eight of the crew had gone
snipe A snipe is any of about 26 wading bird species in three genera in the family Scolopacidae. They are characterized by a very long, slender bill, eyes placed high on the head, and cryptic/ camouflaging plumage. The '' Gallinago'' snipes have ...
shooting in the cutter, with an overnight stay at the Macacu River near Rio. After their return on 2 May, some fell ill with fever. The ship set off on 10 May, a seaman died en route, a ship's boy and a young midshipman died at Bahia. The ship returned to Rio on 3 June. Having confirmed that his measurements were correct, FitzRoy sent corrections to Roussin. At the cottage, Darwin composed his first letter outlining his collecting to Henslow. He said he would not "send a box till we arrive at Monte Video.—it is too great a loss of time both for Carpenters & myself to pack up whilst in harbor". He returned to the ship on 26 June, and they set sail on 5 July. Amidst political changes, ''Beagle'' had a diplomatic role. As they arrived at
Montevideo Montevideo () is the capital and largest city of Uruguay. According to the 2011 census, the city proper has a population of 1,319,108 (about one-third of the country's total population) in an area of . Montevideo is situated on the southern co ...
on 26 July, HMS ''Druid'' signalled them to "clear for action" as British property had been seized in growing unrest after "military usurpation" deposed
Lavalleja Lavalleja () is a department of Uruguay. Its capital is Minas. It is located in the southeast of the country, bordered to the north by the department of Treinta y Tres to the east with Rocha, to the south with Canelones and Maldonado, and to ...
. They took observations for the chronometers, then on 31 July sailed to
Buenos Aires Buenos Aires ( or ; ), officially the Autonomous City of Buenos Aires ( es, link=no, Ciudad Autónoma de Buenos Aires), is the capital and primate city of Argentina. The city is located on the western shore of the Río de la Plata, on South ...
to meet the governor and get maps, but were met by warning shots from a guard ship. FitzRoy promptly lodged a complaint and departed, threatening a
broadside Broadside or broadsides may refer to: Naval * Broadside (naval), terminology for the side of a ship, the battery of cannon on one side of a warship, or their near simultaneous fire on naval warfare Printing and literature * Broadside (comic ...
in response to any further provocation. When they got back on 4 August, FitzRoy informed the ''Druid''s captain who set off to demand an apology. On 5 August, Town officials and the British Consul asked FitzRoy for help to quell a mutiny; the garrison was held by Black troops loyal to Lavalleja. With Darwin and 50 well-armed men from the ship he arrived at the fort, then next day withdrew leaving a stand-off. Darwin enjoyed the excitement, and wrote "It was something new to me to walk with Pistols & Cutlass through the streets of a Town". ''Druid'' returned on 15 August, with a long apology from the government and news that the guard-ship captain had been arrested. Darwin's first box of specimens was ready, and went on the Falmouth packet ''Emulous'' departing on 19 August, Henslow received the box in mid January. On 22 August, after taking soundings in
Samborombón Bay Samborombón Bay () is a bay on the coast of Buenos Aires Province, Argentina. Located at the Río de la Plata's mouth on the Argentine Sea, it begins about southeast of Buenos Aires and is about wide. Toponymy The bay is thought to have been n ...
, ''Beagle'' began survey work down the coast from Cape San Antonio, Buenos Aires Province, Argentina.


Fossil finds

At Bahía Blanca, in the southern part of present
Buenos Aires Province Buenos Aires (), officially the Buenos Aires Province (''Provincia de Buenos Aires'' ), is the largest and most populous Argentine province. It takes its name from the city of Buenos Aires, the capital of the country, which used to be part of th ...
, Darwin rode inland into Patagonia with gauchos: he saw them use bolas to bring down "ostriches" (
rheas The rheas ( ), also known as ñandus ( ) or South American ostriches, are large ratites (flightless birds without a keel on their sternum bone) in the order Rheiformes, native to South America, distantly related to the ostrich and emu. Most tax ...
) and ate roast armadillo. With FitzRoy, he went for "a very pleasant cruize about the bay" on 22 September, and about from the ship, they stopped for a while at Punta Alta. In low cliffs near the point, Darwin found conglomerate rocks containing numerous shells and fossilised teeth and bones of gigantic extinct mammals, in strata near an earth layer with shells and armadillo fossils, suggesting to him quiet tidal deposits rather than a catastrophe.'Cinnamon and port wine': an introduction to the ''Rio Notebook''
Bahía Blanca, September—October 1832.
With assistance (possibly from the young sailor
Syms Covington Syms Covington (1816–1861) was a fiddler and cabin boy on HMS ''Beagle'' who became an assistant to Charles Darwin and was appointed as his personal servant in 1833, continuing in Darwin's service after the voyage until 1839. Originally named ...
acting as his servant), Darwin collected numerous fossils over several days, amusing others with "the cargoes of apparent rubbish which he frequently brought on board". Much of the second day was taken up with excavating a large skull which Darwin found embedded in soft rock, and seemed to him to be allied to the
rhinoceros A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct specie ...
. On 8 October, he returned to the site and found a jawbone and tooth which he was able to identify using
Bory de Saint-Vincent Jean-Baptiste Geneviève Marcellin Bory de Saint-Vincent was a French naturalist, officer and politician. He was born on 6 July 1778 in Agen (Lot-et-Garonne) and died on 22 December 1846 in Paris. Biologist and geographer, he was particularly in ...
's ''Dictionnaire classique''. He wrote home describing this and the large skull as '' Megatherium'' fossils, or perhaps ''
Megalonyx ''Megalonyx'' ( Greek, "large claw") is an extinct genus of ground sloths of the family Megalonychidae, native to North America during the Pliocene and Pleistocene epochs. It became extinct during the Quaternary extinction event at the end ...
'', and excitedly noted that the only specimens in Europe were locked away in the King's collection at
Madrid Madrid ( , ) is the capital and most populous city of Spain. The city has almost 3.4 million inhabitants and a metropolitan area population of approximately 6.7 million. It is the second-largest city in the European Union (EU), and ...
. In the same layer he found a large surface of polygonal plates of bony armour. His immediate thought was that they came from an enormous armadillo like the small creatures common in the area. However, from
Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in nat ...
's misleading description of the Madrid specimen and a recent newspaper report about a fossil collected by
Woodbine Parish Sir Woodbine Parish KCH (14 September 1796, London – 16 August 1882, St. Leonards, Sussex) was a British diplomat, traveller and scientist. The son of Woodbine Parish, of Bawburgh Old Hall, Norfolk, a major in the Light Horse Volunteers, ...
, Darwin thought that the bony armour identified the fossil as ''Megatherium''. With FitzRoy, Darwin went about across the bay to
Monte Hermoso Monte Hermoso is a town located on the Atlantic coast of Argentina, some east of the city of Bahía Blanca, in the south of the Province of Buenos Aires. It is the administrative seat of the partido of Monte Hermoso. Founded at the beginning o ...
on 19 October and found numerous fossils of smaller rodents in contrast to the huge Edentatal mammals of Punta Alta. They returned to Montevideo, and on 2 November revisited Buenos Aires, passing the guard-ship which now gave them due respect. From questioning the finder of the ''Megatherium'' reported in the newspaper (Woodbine Parish’s agent), Darwin concluded it came from the same
geological formation A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exp ...
as his own fossil finds. He also "purchased fragments of some enormous bones" which he "was assured belonged to the former giants!!" In Montevideo from 14 November, he packaged his specimens, including all the fossils, and sent this cargo on the ''Duke of York''
Falmouth packet Packet Newspapers (Cornwall) Limited is a wholly owned subsidiary of the Newsquest media group, which publishes the ''Packet'' series of weekly tabloid newspapers. The series is named after the Falmouth Packet service, which commenced operatio ...
. The mail from home included a copy of the second volume of Charles Lyell's ''Principles of Geology'', a refutation of
Lamarckism Lamarckism, also known as Lamarckian inheritance or neo-Lamarckism, is the notion that an organism can pass on to its offspring physical characteristics that the parent organism acquired through use or disuse during its lifetime. It is also calle ...
in which there was no shared ancestry of different species or overall progress to match the gradual geological change. Instead, it was a continuing cycle in which species mysteriously appeared, closely adapted to their "centres of creation", then became extinct when the environment changed to their disadvantage.


Tierra del Fuego

They reached Tierra del Fuego on 18 December 1832, and Darwin was taken by surprise at what he perceived as the crude savagery of the
Yaghan Yaghan, Yagán or Yahgan may refer to: * Yahgan people, an ethnic group of Argentina and Chile * Yahgan language, their language * Yaghan (dog), an extinct domesticated fox See also

* Yagan (disambiguation) * Yagha, a province of Burkina Faso ...
natives, in stark contrast to the "civilised" behaviour of the three Fuegians they were returning as missionaries (who had been given the names York Minster, Fuegia Basket and
Jemmy Button Orundellico, known as "Jeremy Button" or "Jemmy Button" (c. 1815–1864), was a member of the Yaghan (or Yámana) people from islands around Tierra del Fuego, in modern Chile and Argentina. He was taken to England by Captain FitzRoy in HMS ''B ...
). He described his first meeting with the native Fuegians as being "without exception the most curious and interesting spectacle I ever beheld: I could not have believed how wide was the difference between savage and civilised man: it is greater than between a wild and domesticated animal, inasmuch as in man there is a greater power of improvement." They appeared like "the representations of Devils on the Stage" as in ''
Der Freischütz ' ( J. 277, Op. 77 ''The Marksman'' or ''The Freeshooter'') is a German opera with spoken dialogue in three acts by Carl Maria von Weber with a libretto by Friedrich Kind, based on a story by Johann August Apel and Friedrich Laun from their 1810 ...
''. In contrast, he said of Jemmy that "It seems yet wonderful to me, when I think over all his many good qualities, that he should have been of the same race, and doubtless partaken of the same character, with the miserable, degraded savages whom we first met here." (Four decades later, he recalled these impressions in '' The Descent of Man'' to support his argument that just as humans had descended from "a lower form", civilised society had arisen by graduations from a more primitive state. He recalled how closely the Fuegians on board ''Beagle'' "resembled us in disposition and in most of our mental faculties.") At the island of "Buttons Land" on 23 January 1833, they set up a mission post with huts, gardens, furniture and crockery. Upon returning nine days later, the possessions had been looted and divided up equally by the natives. Matthews gave up, rejoining the ship and leaving the three civilised Fuegians to continue the missionary work. ''Beagle'' went on to the
Falkland Islands The Falkland Islands (; es, Islas Malvinas, link=no ) is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and about from Cape Dubouze ...
, arriving just after the British return. Darwin studied the relationships of species to habitats and found ancient fossils like those he found in Wales. FitzRoy bought a
schooner A schooner () is a type of sailing vessel defined by its rig: fore-and-aft rigged on all of two or more masts and, in the case of a two-masted schooner, the foremast generally being shorter than the mainmast. A common variant, the topsail schoo ...
to assist with the surveying, and they returned to Patagonia, where it was fitted with a new copper bottom and renamed ''Adventure''.
Syms Covington Syms Covington (1816–1861) was a fiddler and cabin boy on HMS ''Beagle'' who became an assistant to Charles Darwin and was appointed as his personal servant in 1833, continuing in Darwin's service after the voyage until 1839. Originally named ...
assisted Darwin in preserving specimens, and his collecting was so successful that, with FitzRoy's agreement, he hired Covington as a full-time servant for £30 a year.


Gauchos, rheas, fossils and geology

The two ships sailed to the Río Negro in Argentina, and on 8 August 1833, Darwin left on another journey inland with the gauchos. On 12 August, he met General Juan Manuel de Rosas who was then leading a punitive expedition in his military campaign against native "Indians" and obtained a passport from him. As they crossed the pampas, the gauchos and Indians told Darwin of a rare smaller species of rhea. After three days at Bahía Blanca, he grew tired of waiting for ''Beagle,'' and on 21 August, revisited Punta Alta where he reviewed the geology of the site in light of his new knowledge, wondering if the bones were older than the seashells. He was very successful with searching for bones, and on 1 September, found a near-complete skeleton with its bones still in position. He set off again and on 1 October, while searching the cliffs of the
Carcarañá River The Carcarañá River (Spanish, Arroyo Saladillo or Río Saladillo) is a river in Argentina. It begins at the confluence of the Río Tercero and the Saladillo River (the lower course of the Río Cuarto) in the south-east of the province of Córd ...
, found "an enormous gnawing tooth", and then, in a cliff of the Paraná River, saw "two great groups of immense bones" which were too soft to collect but a tooth fragment identified them as
mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...
s. Illness delayed him at Santa Fe, and after seeing the fossilised casing of a huge armadillo embedded in rock, he was puzzled to find a horse tooth in the same rock layer since horses had been introduced to the continent with European migration. They took a riverboat down the Paraná River to
Buenos Aires Buenos Aires ( or ; ), officially the Autonomous City of Buenos Aires ( es, link=no, Ciudad Autónoma de Buenos Aires), is the capital and primate city of Argentina. The city is located on the western shore of the Río de la Plata, on South ...
but became entangled in a revolution as rebels allied to Rosas blockaded the city. The passport helped, and with Covington, he managed to escape in a boatload of refugees. They rejoined ''Beagle'' at Montevideo. As surveys were still in progress, Darwin set off on another "galloping" trip in Banda Oriental to see the Uruguay River and visit the Estancia of Mr Keen near Mercedes on the Río Negro. On 25 November, he "heard of some giants bones, which as usual turned out to be those of the Megatherium" but could only extract a few broken fragments. The next day, he visited a nearby house and bought "a head of a Megatherium which must have been when found quite perfect" for about two
shilling The shilling is a historical coin, and the name of a unit of modern currencies formerly used in the United Kingdom, Australia, New Zealand, other British Commonwealth countries and Ireland, where they were generally equivalent to 12 pence ...
s, though the teeth had since been broken and the lower jaw had been lost. Mr Keen arranged to ship the skull downriver to Buenos Aires. At Las Piedras, a clergyman let him see fossils, including a club-like tail which he sketched and called an "extraordinary weapon".'Banda Oriental S. Cruz.' Beagle field notebook. EH1.9
p. 36
a typical Glyptodont tail.
His notes included a page showing his realisation that the cliff banks of the rivers exposed two strata formed in an
estuary An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
interrupted by an undersea stratum, indicating that the land had risen and fallen.'Banda Oriental S. Cruz.' Beagle field notebook. EH1.9
p. 37
/ref> Back at Montevideo, Darwin was introduced to Conrad Martens, the replacement artist brought on board ''Beagle'' after Augustus Earle had to leave due to health problems. They sailed south, putting in at Port Desire on 23 December, and the following day Darwin shot a guanaco which provided them with a Christmas meal. Early in the new year, Martens shot a rhea which they enjoyed eating before Darwin realised that this was the elusive smaller rhea and preserved the remains. On 9 January 1834, further south, they reached
Port St Julian A port is a maritime facility comprising one or more wharves or loading areas, where ships load and discharge cargo and passengers. Although usually situated on a sea coast or estuary, ports can also be found far inland, such as Ham ...
and exploring the local geology in cliffs near the harbour Darwin found fossils of pieces of spine and a hind leg of "some large animal, I fancy a Mastodon". On 26 January, they entered the Straits of Magellan, and at St. Gregory's Bay, they met half-civilised Patagonian "giants" over tall, described by Darwin as "excellent practical naturalists". One told him that the smaller rheas were the only species this far south, while the larger rheas kept to the north, the species meeting around the Rio Negro. After further surveying in Tierra del Fuego, they returned on 5 March 1834 to visit the missionaries but found the huts deserted. Then canoes approached, and they found that one of the natives was Jemmy Button, who had lost his possessions and had settled into the native ways, taking a wife. Darwin had never seen "so complete & grievous a change". Jemmy came on board and dined using his cutlery properly, speaking English as well as ever, then assured them that he "had not the least wish to return to England" and was "happy and contented", leaving them gifts of otter skins and arrowheads before returning to the canoe to join his wife. Of the first visit, Darwin had written that "Viewing such men, one can hardly make oneself believe that they are fellow creatures placed in the same world. .... It is a common subject of conjecture; what pleasure in life some of the less gifted animals can enjoy? How much more reasonably it may be asked with respect to these men", yet Jemmy had readily adapted to civilisation and then chosen to return to his primitive ways. This raised awkward questions; it jarred with Charles Lyell's sheltered views, expressed in volume 2 of his ''Principles of Geology'', that human races "showed only a slight deviation from a common standard", and that acceptance of transmutation meant renouncing man's "belief in the high genealogy of his species". About this time Darwin wrote ''Reflection on Reading My Geological Notes'', the first of a series of essays included in his notes. He speculated on possible causes of the land repeatedly being raised, and on a history of life in Patagonia as a sequence of named species. They returned to the Falkland Islands on 16 March, just after an incident where gauchos and Indians had butchered senior members of Vernet's settlement, and helped to put the revolt down. Darwin noted the immense number of organisms dependent on the kelp forests. He received word from Henslow that his first dispatch of fossils had reached Cambridge, were highly prized by the expert William Clift as showing hitherto unknown species and features of the ''Megatherium'', and had been displayed by William Buckland and Clift before the cream of British science, making Darwin's reputation. ''Beagle'' now sailed to southern Patagonia, and on 19 April, an expedition including FitzRoy and Darwin set off to take boats as far as possible up the Santa Cruz river, with all involved taking turns in teams dragging the boats upstream. The river cut through a series of rises, then through plateaux forming wide plains covered with shells and shingle. Darwin discussed with FitzRoy his interpretation that these terraces had been shores that had gradually raised per Lyell's theories. Several of the smaller rheas were seen in the distance but were too elusive to catch. The expedition approached the Andes but had to turn back. Darwin summarised his speculation in his essay on the ''Elevation of Patagonia''. Though tentative, it challenged Lyell's ideas. Darwin drew on measurements by ''Beagle'' officers, as well as his own measurements, to propose that the plains had been raised in successive stages by forces acting over a wide area, rather than smaller-scale actions in a continuous movement. However, he supported Lyell in finding evidence to dismiss a sudden deluge when normal processes were suddenly speeded. Seashells he had found far inland still showing their colour suggested to him that the process had been relatively recent and could have affected human history.


West coast of South America

''Beagle'' and ''Adventure'' now surveyed the Straits of Magellan before sailing north up the west coast, reaching
Chiloé Island Chiloé Island ( es, Isla de Chiloé, , ) also known as Greater Island of Chiloé (''Isla Grande de Chiloé''), is the largest island of the Chiloé Archipelago off the west coast of Chile, in the Pacific Ocean. The island is located in southern ...
in the wet and heavily wooded Chiloé Archipelago on 28 June 1834. They then spent the next six months surveying the coast and islands southwards. On Chiloé, Darwin found fragments of black
lignite Lignite, often referred to as brown coal, is a soft, brown, combustible, sedimentary rock formed from naturally compressed peat. It has a carbon content around 25–35%, and is considered the lowest rank of coal due to its relatively low heat ...
and
petrified In geology, petrifaction or petrification () is the process by which organic material becomes a fossil through the replacement of the original material and the filling of the original pore spaces with minerals. Petrified wood typifies this p ...
wood, at least two of which the British Geological Survey discovered in 2011 locked away in their collection labelled "unregistered fossil plants". Exchanged with Joseph Dalton Hooker about ten years later, one slide was signed "Chiloe, C. Darwin Esq". They arrived at
Valparaíso Valparaíso (; ) is a major city, seaport, naval base, and educational centre in the commune of Valparaíso, Chile. "Greater Valparaíso" is the second largest metropolitan area in the country. Valparaíso is located about northwest of Santiago ...
on 23 July, and Darwin "got scent of some fossil bones .... if gold or galloping will get them, they shall be mine." After several walks in the area, he obtained horses and, on 14 August, set off up the volcanic
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
with a companion. Three days later they spent an enjoyable day on the summit of the "Campana or Bell" mountain,
Cerro La Campana Cerro la Campana, the Bell mountain, is a mountain in La Campana National Park in central Chile. The Pacific and the mountain Aconcagua are visible from the summit on clear days. Due to the area's expanding human population, considerable def ...
. Darwin visited a copper mine and spent five days scrambling in the mountains before going on to Santiago, Chile. On his way back, he fell ill on 20 September and had to spend a month in bed. It is possible that he contracted Chagas' disease here, leading to his health problems after his return to England, but this diagnosis of his symptoms is disputed. He learnt that the Admiralty had reprimanded FitzRoy for buying ''Adventure''. FitzRoy had taken it badly, selling the ship and announcing they would go back to recheck his survey. He then had resigned his command, doubting his sanity, but was persuaded by his officers to withdraw his resignation and proceed. The artist Conrad Martens left the ship and took passage to Australia. After waiting for Darwin, ''Beagle'' sailed on 11 November to survey the Chonos Archipelago. From here, they saw the eruption of the volcano Osorno in the Andes. They sailed north, and Darwin wondered about the fossils he had found. The giant ''
Mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...
''s and ''Megatherium''s were extinct, but he had found no geological signs of a " diluvial debacle" or of the changed circumstances that, in Lyell's view, led to species no longer being adapted to the position they were created to fit. He agreed with Lyell's idea of "the gradual birth & death of species", but, unlike Lyell, Darwin was willing to believe Giovanni Battista Brocchi's idea that extinct species had somehow aged and died out. They arrived at the port of
Valdivia Valdivia (; Mapuche: Ainil) is a city and commune in southern Chile, administered by the Municipality of Valdivia. The city is named after its founder Pedro de Valdivia and is located at the confluence of the Calle-Calle, Valdivia, and Cau-Ca ...
on 8 February 1835. Twelve days later, Darwin was on shore when he experienced a severe earthquake and returned to find the port town badly damaged. They sailed north to Concepción, and arrived on 4 March to find that the same earthquake had devastated the city by repeated shocks and a tsunami, with even the cathedral in ruins. Darwin noted the horrors of death and destruction, and FitzRoy carefully established that mussel beds were now above high tide, giving clear evidence of the ground rising some , which he confirmed a month later. They had actually experienced the gradual process of the continent emerging from the ocean, as Lyell had indicated. They returned to Valparaiso on 11 March, Darwin set out on another trek up the Andes three days later and, on 21 March, reached the continental divide at : even here, he found fossil seashells in the rocks. He felt the glorious view "was like watching a thunderstorm, or hearing in the full Orchestra a Chorus of the ''Messiah''." After going on to Mendoza, they were returning by a different pass when they found a petrified forest of fossilised trees, crystallised in a sandstone escarpment showing him that they had been on a Pacific beach when the land sank, burying them in the sand which had been compressed into rock, then had gradually been raised with the continent to stand at in the mountains. On returning to Valparaiso with half a mule's load of specimens, he wrote to his family on 23 April that his findings, if accepted, would be crucial to the theory of the formation of the world. After another gruelling expedition in the Andes, while ''Beagle'' was refitted, he rejoined it at
Copiapó Copiapó () is a city and commune in northern Chile, located about 65 kilometers east of the coastal town of Caldera. Founded on December 8, 1744, it is the capital of Copiapó Province and Atacama Region. Copiapó lies about 800 km nor ...
on 5 July and sailed to
Lima Lima ( ; ), originally founded as Ciudad de Los Reyes (City of The Kings) is the capital and the largest city of Peru. It is located in the valleys of the Chillón, Rímac and Lurín Rivers, in the desert zone of the central coastal part of ...
but found an armed insurrection in progress and had to stay with the ship. Here he was writing up his notes when he realised that Lyell's idea, that coral atolls were on the rims of rising extinct volcanoes, made less sense than the volcanoes gradually sinking so that the coral reefs around the island kept building themselves close to sea level and became an atoll as the volcano disappeared below. This was a theory he would examine when they reached such islands. On 14 June, when about to leave Valparaiso, FitzRoy had received news of the shipwreck of HMS ''Challenger'' captained by his friend Michael Seymour (Darwin had arranged two boxes for this packet ship early in the year). On investigation, FitzRoy found that Commodore Mason was unwilling to take to the rescue for fear of lee-shore hazards, so FitzRoy "had to bully him & at last offered to go as Pilot". After "a tremendous quarrel" with hints to the Commodore of court-martial, they took ''Blonde'' to Concepción. FitzRoy rode about on horseback with a guide to reach Seymour's camp at the Lebu River, then returned to further disputes before ''Blonde'' set out and rescued the survivors of the shipwreck on 5 July. Wickham took ''Beagle'' on to reach Copiapò on 3 July, two days before Darwin rejoined the ship and they continued on to Lima. On 9 September ''Blonde'' brought FitzRoy to join them at Lima.


Galápagos Islands

A week out of Lima, ''Beagle'' reached the
Galápagos Islands The Galápagos Islands ( Spanish: , , ) are an archipelago of volcanic islands. They are distributed on each side of the equator in the Pacific Ocean, surrounding the centre of the Western Hemisphere, and are part of the Republic of Ecuad ...
on 15 September 1835. The next day Captain FitzRoy dropped anchor near where the town of
Puerto Baquerizo Moreno Puerto Baquerizo Moreno () is the Capital (political), capital of Galápagos Province, Ecuador. It is located on the southwestern coast of San Cristóbal (Galápagos Islands), San Cristóbal, the easternmost island in the archipelago, and is the c ...
is now sited, at Chatham Island. At the location that is now known as Frigatebird Hill (Cerro Tijeretas), Darwin spent his first hour onshore in the Galapagos islands. Darwin eagerly looked forward to seeing newly formed volcanic islands and took every opportunity to go ashore while ''Beagle'' was methodically moved round to chart the coast. He found broken black rocky volcanic lava scorching under the hot sun, and made detailed geological notes of features including volcanic cones like chimneys which reminded him of the iron foundries of industrial
Staffordshire Staffordshire (; postal abbreviation Staffs.) is a landlocked county in the West Midlands region of England. It borders Cheshire to the northwest, Derbyshire and Leicestershire to the east, Warwickshire to the southeast, the West Midlands C ...
. He was disappointed that he did not see active volcanoes or find strata showing uplift as he had hoped, though one of the officers found broken oyster-shells high above the sea on one of the islands. Abundant giant
Galápagos tortoise The Galápagos tortoise or Galápagos giant tortoise (''Chelonoidis niger'') is a species of very large tortoise in the genus ''Chelonoidis'' (which also contains three smaller species from mainland South America). It comprises 15 subspecies ...
s appeared to him almost
antediluvian The antediluvian (alternatively pre-diluvian or pre-flood) period is the time period chronicled in the Bible between the fall of man and the Genesis flood narrative in biblical cosmology. The term was coined by Thomas Browne. The narrative tak ...
, and large black marine iguanas seemed "most disgusting, clumsy Lizards" well suited to their habitat—he noted that someone had called them "imps of darkness". Darwin had learnt from Henslow about studying the geographical distribution of species, and particularly of linked species on oceanic islands and nearby continents, so he endeavoured to collect plants in flower. He found widespread "wretched-looking" thin scrub thickets of only ten species and very few insects. Birds were remarkably unafraid of humans, and in his first field note, he recorded that a mockingbird was similar to those he had seen on the continent. ''Beagle'' sailed on to Charles Island. By chance, they were greeted by the "Englishman"
Nicholas Lawson Nicholas Oliver Lawson (born Nicolai Olaus Lossius; 23 November 1790 – 1 March 1851) was a Norwegian-born, vice governor of Galápagos for the Republic of Ecuador. While there, he provided information which contributed to Charles Darwin's first ...
, acting Governor of Galápagos for the Republic of the Equator, who accompanied them up to the penal colony. It was said that tortoises differed in the shape of the shells from island to island, and Darwin noted Lawson's statement that on seeing a tortoise, he could "pronounce with certainty from which island it has been brought". Though Darwin remembered this later, he did not pay much attention at the time. However, he found a mockingbird and "fortunately happened to observe" that it differed from the Chatham Island specimen, so from then on, he carefully noted where mockingbirds had been caught. He industriously collected all the animals, plants, insects and reptiles, and speculated about finding "from future comparison to what district or 'centre of creation' the organized beings of this archipelago must be attached." At this stage, his thoughts reflected Lyell's rejection of
transmutation of species Transmutation of species and transformism are unproven 18th and 19th-century evolutionary ideas about the change of one species into another that preceded Charles Darwin's theory of natural selection. The French ''Transformisme'' was a term used ...
. They went on to Albemarle Island, where Darwin saw a small jet of smoke from a recently active volcano. On 1 October, he landed near Tagus Cove and explored Beagle Crater. There, he saw his first
Galapagos land iguana The Galápagos land iguana (''Conolophus subcristatus'') is a very large species of lizard in the family Iguanidae. It is one of three species of the genus ''Conolophus''. It is endemic to the Galápagos Islands, in the dry lowlands of the island ...
s. Water pits were disappointingly inadequate for drinking, but attracted swarms of small birds, and Darwin made his only note of the finches he was not bothering to label by island. He caught a third species of mockingbird. After passing the northern islands of Abingdon,
Tower A tower is a tall structure, taller than it is wide, often by a significant factor. Towers are distinguished from masts by their lack of guy-wires and are therefore, along with tall buildings, self-supporting structures. Towers are specific ...
and Bindloe, Darwin was put ashore at James Island for nine days together with the surgeon Benjamin Bynoe and their servants. They busily collected all sorts of specimens while ''Beagle'' went back to Chatham Island for freshwater. After further surveying, ''Beagle'' set sail for Tahiti on 20 October 1835. Darwin wrote up his notes, and to his astonishment, found that all the mockingbirds caught on Charles, Albemarle, James and Chatham Islands differed from island to island. He wrote "This birds which is so closely allied to the Thenca of Chili (Callandra of B. Ayres) is singular from existing as varieties or distinct species in the different Isds.— I have four specimens from as many Isds.— These will be found to be 2 or 3 varieties.— Each variety is constant in its own Island....".


Tahiti to Australia

They sailed on, dining on Galapagos tortoises, and passed the
atoll An atoll () is a ring-shaped island, including a coral rim that encircles a lagoon partially or completely. There may be coral islands or cays on the rim. Atolls are located in warm tropical or subtropical oceans and seas where corals can gro ...
of Honden Island on 9 November. They passed through the
Low Islands The Low Islands are one of the many uninhabited Canadian arctic island groups in Qikiqtaaluk Region, Nunavut. They are located in Ungava Bay, stretching between Young Island to the south and Lookout Island to the north. References Extern ...
archipelago, with Darwin remarking that they had "a very uninteresting appearance; a long brilliantly white beach is capped by a low bright line of green vegetation." Arriving at Tahiti on 15 November, he soon found interest in luxuriant vegetation and the pleasant intelligent natives who showed the benefits of Christianity, refuting allegations he had read about tyrannical missionaries overturning indigenous cultures. On 19 December, they reached New Zealand, where Darwin thought the tattooed
Māori Māori or Maori can refer to: Relating to the Māori people * Māori people of New Zealand, or members of that group * Māori language, the language of the Māori people of New Zealand * Māori culture * Cook Islanders, the Māori people of the Co ...
to be savages with the character of a much lower order than the Tahitians. He also noted that they and their homes were "filthily dirty and offensive". Darwin saw missionaries bringing improvement in character, as well as new farming practices with an exemplary "English farm" employing natives. Richard Matthews was left here with his elder brother Joseph Matthews who was a missionary at Kaitaia. Darwin and FitzRoy were agreed that missionaries had been unfairly misrepresented in tracts, particularly one written by the artist
Augustus Earle Augustus Earle (1793–1838) was a British painter. Unlike earlier artists who worked outside Europe and were employed on voyages of exploration or worked abroad for wealthy, often aristocratic patrons, Earle was able to operate quite indepe ...
which he had left on the ship. Darwin also noted many English residents of the most worthless character, including runaway convicts from
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
. By 30 December, he was glad to leave New Zealand. The first sight of Australia on 12 January 1836 reminded him of Patagonia, but inland the country improved, and he was soon filled with admiration at the bustling city of
Sydney Sydney ( ) is the capital city of the state of New South Wales, and the most populous city in both Australia and Oceania. Located on Australia's east coast, the metropolis surrounds Sydney Harbour and extends about towards the Blue Mounta ...
. On a journey into the interior, he came across a group of Aboriginal peoples who looked "good-humoured & pleasant & they appeared far from such utterly degraded beings as usually represented". They gave him a display of spear throwing for a shilling, and he reflected sadly on how their numbers were rapidly decreasing. At a large sheep farm, he joined a hunting party and caught his first marsupial, a "
potoroo Potoroo is a common name for species of ''Potorous'', a genus of smaller marsupials. They are allied to the Macropodiformes, the suborder of kangaroo, wallaby, and other rat-kangaroo genera. All three extant species are threatened by ecological ...
" (rat-kangaroo). Reflecting on the strange animals of the country, he thought that an unbeliever "might exclaim 'Surely two distinct Creators must have been twork; their object however has been the same & certainly the end in each case is complete'," yet an
antlion The antlions are a group of about 2,000 species of insect in the neuropteran family Myrmeleontidae. They are known for the predatory habits of their larvae, which mostly dig pits to trap passing ants or other prey. In North America, the larvae ...
he was watching was very similar to its European counterpart. That evening he saw the even stranger
platypus The platypus (''Ornithorhynchus anatinus''), sometimes referred to as the duck-billed platypus, is a semiaquatic, egg-laying mammal endemic to eastern Australia, including Tasmania. The platypus is the sole living representative or mono ...
and noticed that its bill was soft, unlike the preserved specimens he had seen. Aboriginal stories that they laid eggs were believed by few Europeans. ''Beagle'' visited
Hobart Hobart ( ; Nuennonne/ Palawa kani: ''nipaluna'') is the capital and most populous city of the Australian island state of Tasmania. Home to almost half of all Tasmanians, it is the least-populated Australian state capital city, and second-small ...
,
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
, where Darwin was impressed by the agreeable high society of the settlers but noted that the island's "Aboriginal blacks are all removed & kept (in reality as prisoners) in a Promontory, the neck of which is guarded. I believe it was not possible to avoid this cruel step; although without doubt the misconduct of the Whites first led to the Necessity." They then sailed to King George's Sound in south-west Australia, a dismal settlement then being replaced by the
Swan River Colony The Swan River Colony, also known as the Swan River Settlement, or just Swan River, was a British colony established in 1829 on the Swan River, in Western Australia. This initial settlement place on the Swan River was soon named Perth, and it ...
. Darwin was impressed by the "good disposition of the aboriginal blacks... Although true Savages, it is impossible not to feel an inclination to like such quiet good-natured men." He provided boiled rice for an aboriginal " Corrobery" dancing party performed by the men of two tribes to the great pleasure of the women and children, a "most rude barbarous scene" in which everyone appeared in high spirits, "all moving in hideous harmony" and "perfectly at their ease". ''Beagle'' departure in a storm was delayed when she ran aground. She was refloated and got on her way.


Keeling Island homewards

FitzRoy's instructions from the Admiralty required a detailed geological survey of a circular
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and se ...
atoll An atoll () is a ring-shaped island, including a coral rim that encircles a lagoon partially or completely. There may be coral islands or cays on the rim. Atolls are located in warm tropical or subtropical oceans and seas where corals can gro ...
to investigate how
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s formed, particularly whether they rose from the bottom of the sea or the summits of extinct volcanoes, and the effects of
tide Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another. Tide tables ...
s measured with specially constructed gauges. He chose the Keeling Islands in the
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by ...
, and on arrival on 1 April, the entire crew set to work. Darwin found a coconut economy, serving both the small settlement and wildlife. There was a limited range of native plants and no land birds, but hermit crabs everywhere. The lagoons teemed with a wide variety of invertebrates and fish, and he examined the atoll's structure in view of the theory he had developed in Lima, of encircling reefs becoming atolls as an island sank. This idea was supported by the numerous soundings FitzRoy had taken, showing a steep slope outside the reef with no living corals below 20–30
fathom A fathom is a unit of length in the imperial and the U.S. customary systems equal to , used especially for measuring the depth of water. The fathom is neither an International Standard (SI) unit, nor an internationally-accepted non-SI unit. ...
s (40–60 m). Arriving at
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label= Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It ...
on 29 April 1836, Darwin was impressed by the civilised prosperity of the French colony, which had come under British rule. He toured the island, examining its volcanic mountains and fringing coral reefs. The Surveyor-general Captain Lloyd took him on the only elephant on the island to see an elevated coral plain. By then, FitzRoy was writing the official ''Narrative'' of the ''Beagle'' voyages, and after reading Darwin's diary he proposed a joint publication. Darwin asked his family about FitzRoy's idea "to have the disposal & arranging of my journal & to mingle it with his own". ''Beagle'' reached the Cape of Good Hope on 31 May. In
Cape Town Cape Town ( af, Kaapstad; , xh, iKapa) is one of South Africa's three capital cities, serving as the seat of the Parliament of South Africa. It is the legislative capital of the country, the oldest city in the country, and the second largest ...
, Darwin received a letter dated 29 January from his sister Catherine which briefly mentioned "the little books, with the Extracts from your Letters; every body is much pleased, with them, who has seen them". Darwin was horrified that his careless words were in print, but ''No hay remedio'' (it can't be helped). Unknown to Darwin, his fame was spreading; extracts from his letters to Henslow had been read to the
Cambridge Philosophical Society The Cambridge Philosophical Society (CPS) is a scientific society at the University of Cambridge. It was founded in 1819. The name derives from the medieval use of the word philosophy to denote any research undertaken outside the fields of l ...
on 16 November 1835 by Henslow and Sedgwick. On 18 November, Sedgwick had read extracts to the Geological Society of London, and this had been reported in '' The Athenæum'' on 21 November. On 25 December, their father received a letter from Henslow which said that Darwin would become one of the premier naturalists of the time and enclosed some copies of the pamphlet '' Extracts from letters addressed to Professor Henslow'' which had been printed for private distribution. Their father "did not move from his seat till he had read every word of book & he was very much gratified – he liked so much the simple clear way you gave your information". Darwin explored the geology of the area, reaching conclusions about the slate formation and the injection of granite seams as a liquid which differed from the ideas of Lyell and Sedgwick. The zoologist Andrew Smith showed him formations, and later discussed the large animals living on sparse vegetation, showing that a lack of luxuriant vegetation did not explain the extinction of the giant creatures in South America. Around 15 June, Darwin and FitzRoy visited the noted astronomer Sir
John Herschel Sir John Frederick William Herschel, 1st Baronet (; 7 March 1792 – 11 May 1871) was an English polymath active as a mathematician, astronomer, chemist, inventor, experimental photographer who invented the blueprint and did botanical wo ...
. In his diary, Darwin called this "the most memorable event which, for a long period, I have had the good fortune to enjoy." His zeal for science had been stirred at Cambridge by reading Herschel's book on philosophy of science, which had guided his theorising during the voyage. Their discussion is not recorded, but a few months earlier, on 20 February 1836, Herschel had written to Lyell praising his ''Principles of Geology'' as a work which would bring "a complete revolution in tssubject, by altering entirely the point of view in which it must thenceforward be contemplated." and opening a way for bold speculation on "that mystery of mysteries, the replacement of extinct species by others." Herschel himself thought catastrophic extinction and renewal "an inadequate conception of the Creator", and by analogy with other intermediate causes "the origination of fresh species, could it ever come under our cognizance, would be found to be a natural in contradistinction to a miraculous process".
In Cape Town, missionaries were being accused of causing racial tension and profiteering, and after ''Beagle'' set to sea on 18 June, FitzRoy wrote an open letter to the
evangelical Evangelicalism (), also called evangelical Christianity or evangelical Protestantism, is a worldwide interdenominational movement within Protestant Christianity that affirms the centrality of being " born again", in which an individual expe ...
''South African Christian Recorder'' on the ''Moral State of Tahiti'' incorporating extracts from both his and Darwin's diaries to defend the reputation of missionaries. This was given to a passing ship that took it to Cape Town to become FitzRoy's (and Darwin's) first published work. On 8 July, they stopped at St. Helena for six days. Darwin took lodgings near
Napoleon Napoleon Bonaparte ; it, Napoleone Bonaparte, ; co, Napulione Buonaparte. (born Napoleone Buonaparte; 15 August 1769 – 5 May 1821), later known by his regnal name Napoleon I, was a French military commander and political leader wh ...
's tomb, and when writing to Henslow asking to be proposed for the Geological Society, he mentioned his suspicions "that differently from most Volcanic Islds. its structure is rather complicated. It seems strange, that this little centre of a distinct creation should, as is asserted, bear marks of recent elevation." With a guide, he wandered over the island, noting its complex sloping strata showing fault lines, interlaced with
volcanic A volcano is a rupture in the crust of a planetary-mass object, such as Earth, that allows hot lava, volcanic ash, and gases to escape from a magma chamber below the surface. On Earth, volcanoes are most often found where tectonic plat ...
dykes. He examined beds high on the hill that had been taken as seashells showing that St. Helena had risen from the ocean in recent times, but Darwin identified them as extinct species of land-shells. He noted that woodland had been destroyed by goats and hogs that had run wild since being introduced in 1502, and native vegetation only predominated on high steep ridges, having been replaced by imported species. At this stage, Darwin had an acute interest in the island
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
, and his description of St Helena as "a little centre of creation" in his geological diary reflects Charles Lyell's speculation in volume 2 of ''Principles of Geology'' that the island would have acted as a "focus of creative force". He later recalled believing in the permanence of species, but "as far as I can remember, vague doubts occasionally flitted across my mind". When organising his ''Ornithological Notes'' between mid June and August, Darwin expanded on his initial notes on the Galapagos mockingbird ''Mimus thenca'':
These birds are closely allied in appearance to the Thenca of Chile or Callandra of la Plata. ... In each Isld. each kind is found: habits of all are indistinguishable. When I recollect, the fact that the form of the body, shape of scales & general size, the Spaniards can at once pronounce, from which Island any Tortoise may have been brought. When I see these Islands in sight of each other, & ut del.possessed of but a scanty stock of animals, tenanted by these birds, but slightly differing in structure & filling the same place in Nature, I must suspect they are only varieties.
The only fact of a similar kind of which I am aware, is the constant asserted difference – between the wolf-like Fox of East & West Falkland Islds.
If there is the slightest foundation for these remarks the zoology of Archipelagoes – will be well worth examining; for such facts ould ''inserted''undermine the stability of Species.
The term "would" before "undermine" had been added after writing what is now noted as the first expression of his doubts about species being immutable. That led to him becoming convinced about the
transmutation of species Transmutation of species and transformism are unproven 18th and 19th-century evolutionary ideas about the change of one species into another that preceded Charles Darwin's theory of natural selection. The French ''Transformisme'' was a term used ...
and hence evolution.
   
In opposing transmutation, Lyell had proposed that varieties arose due to changes in the environment, but these varieties lived in similar conditions though each on its own island. Darwin had just reviewed similar inconsistencies with mainland bird genera such as ''
Pteroptochos ''Pteroptochos'' is a genus of bird in the family Rhinocryptidae.Chesser RT (1999Molecular Systematics of the Rhinocryptid Genus ''Pteroptochos'' ''The Condor'' 101(2): 439–446. https://doi.org/10.2307/1370012 Species It contains the follow ...
''. Though his suspicions about the Falkland Island fox may have been unsupported, the differences in
Galápagos tortoise The Galápagos tortoise or Galápagos giant tortoise (''Chelonoidis niger'') is a species of very large tortoise in the genus ''Chelonoidis'' (which also contains three smaller species from mainland South America). It comprises 15 subspecies ...
s between islands were remembered, and he later wrote that he had been greatly struck from around March 1836 by the character of South American fossils and species on the Galapagos Archipelago, noting "These facts origin (especially latter) of all my views". ''Beagle'' reached
Ascension Island Ascension Island is an isolated volcanic island, 7°56′ south of the Equator in the South Atlantic Ocean. It is about from the coast of Africa and from the coast of South America. It is governed as part of the British Overseas Territory of ...
on 19 July 1836, and Darwin was delighted to receive letters from his sisters with news that Sedgwick had written to Dr. Butler: "He is doing admirably in S. America, & has already sent home a Collection above all praise.— It was the best thing in the world for him that he went out on the Voyage of Discovery— There was some risk of his turning out an idle man: but his character will now be fixed, & if God spare his life, he will have a great name among the Naturalists of Europe." Darwin later recalled how he "clambered over the mountains... with a bounding step and made the volcanic rocks resound under my geological hammer!." He agreed with the saying attributed to the people of St Helena that "We know we live on a rock, but the poor people at Ascension live on a cinder", and noted the care taken to sustain "houses, gardens & fields placed near the summit of the central mountain". (In the 1840s, Darwin worked with Hooker, who proposed in 1847 that the Royal Navy shall import tree species, a project started in 1850 which led to the creation of an artificial cloud forest on what is now
Green Mountain Green Mountain is a common name for "The Peak", the highest point on Ascension Island, which has gained some fame for claims that it is one of very few large-scale artificial forests. History and vegetation Many early 19th-century accounts, inc ...
.) On 23 July, they set off again longing to reach home, but FitzRoy, who wanted to ensure the accuracy of his longitude measurements, took the ship across the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
back to Bahia in Brazil to take check readings. Darwin was glad to see the beauties of the jungle for one last time but now compared "the stately Mango trees with the Horse Chesnuts of England." The return trip was delayed for a further 11 days when weather forced ''Beagle'' to shelter further up the coast at
Pernambuco Pernambuco () is a state of Brazil, located in the Northeast region of the country. With an estimated population of 9.6 million people as of 2020, making it seventh-most populous state of Brazil and with around 98,148 km², being the ...
, where Darwin examined rocks for signs of elevation, noted "Mangroves like rank grass", and investigated marine invertebrates at various depths on the sandbar. ''Beagle'' departed for home on 17 August. After a stormy passage including a stop for supplies at the
Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...
, the Beagle finally reached England on 2 October 1836 and anchored at Falmouth, Cornwall.


Return

On the stormy night of 2 October 1836, immediately after arriving in Falmouth, Darwin set off on the
Royal Mail , kw, Postya Riel, ga, An Post Ríoga , logo = Royal Mail.svg , logo_size = 250px , type = Public limited company , traded_as = , foundation = , founder = Henry VIII , location = London, England, UK , key_people = * Keith Williams ...
coach Coach may refer to: Guidance/instruction * Coach (sport), a director of athletes' training and activities * Coaching, the practice of guiding an individual through a process ** Acting coach, a teacher who trains performers Transportation * Coac ...
from Fish Strand Hill (a plaque now marks the site – see photographs.) for the two day journey to his family home, The Mount House in Shrewsbury, Shropshire. As he wrote to FitzRoy, the countryside they passed was "beautiful & cheerful", and though the "stupid people on the coach did not seem to think the fields one bit greener than usual", he now knew "that the wide world does not contain so happy a prospect as the rich cultivated land of England". In April 1835 Darwin had written that he was undecided whether to "sleep at the Lion, the first night," when he arrived by the daily Wonder coach, "or disturb you all in the dead of the night". He arrived late at night on 4 October 1836, and at breakfast the next morning greeted his family, to their delight and celebrations. His sisters assured him he did "not look the least different", but his father's first reaction was to tell them "Why, the shape of his head is quite altered." After time spent catching up with family, Darwin went on to
Cambridge Cambridge ( ) is a university city and the county town in Cambridgeshire, England. It is located on the River Cam approximately north of London. As of the 2021 United Kingdom census, the population of Cambridge was 145,700. Cambridge bec ...
on 15 October and sought Henslow's advice on organising the description and cataloguing of his collections. Darwin's father gave him an allowance that enabled him to put aside other careers. As a scientific celebrity with a reputation established by his fossils and the wide distribution of '' Extracts from Letters to Henslow'' on South American natural history and geology, Darwin toured
London London is the capital and List of urban areas in the United Kingdom, largest city of England and the United Kingdom, with a population of just under 9 million. It stands on the River Thames in south-east England at the head of a estuary dow ...
's social institutions. By this time, he was part of the "scientific establishment", collaborating with expert naturalists to describe his specimens and working on ideas he had been developing during the voyage. Charles Lyell gave him enthusiastic backing. In December 1836, Darwin presented a talk to the
Cambridge Philosophical Society The Cambridge Philosophical Society (CPS) is a scientific society at the University of Cambridge. It was founded in 1819. The name derives from the medieval use of the word philosophy to denote any research undertaken outside the fields of l ...
. He wrote a paper proving that Chile, and the South American continent, was slowly rising, which he read to the Geological Society of London on 4 January 1837. Darwin was willing to have his diary published mixed in with FitzRoy's account, but his relatives, including Emma and
Hensleigh Wedgwood Hensleigh Wedgwood (21 January 1803 – 2 June 1891) was a British etymologist, philologist and barrister, author of '' A Dictionary of English Etymology''. He was a cousin of Charles Darwin, whom his sister Emma married in 1839. Early life We ...
, urged that it be published separately. On 30 December, the question was settled by FitzRoy taking the advice of
William Broderip William John Broderip FRS (21 November 1789 – 27 February 1859) was an English lawyer and naturalist. Life Broderip, the eldest son of William Broderip, surgeon from Bristol, was born at Bristol on 21 November 1789, and, after being educat ...
that Darwin's journal should form the third volume of the ''Narrative''. Darwin set to work reorganising his diary, trimming it, and incorporating scientific material from his notes. He completed his '' Journal and Remarks'' (now commonly known as ''The Voyage of the Beagle'') in August 1837, but FitzRoy was slower, and the three volumes were published in August 1839.
Syms Covington Syms Covington (1816–1861) was a fiddler and cabin boy on HMS ''Beagle'' who became an assistant to Charles Darwin and was appointed as his personal servant in 1833, continuing in Darwin's service after the voyage until 1839. Originally named ...
stayed with Darwin as his servant. Then, on 25 February 1839 (shortly after Darwin's marriage), Covington left on good terms and migrated to Australia.


Expert publications on Darwin's collections

Darwin had shown great ability as a collector and had done the best he could with the reference books he had on the ship. It was now the province of recognised expert specialists to establish which specimens were unknown, and make their considered taxonomic decisions on defining and naming new species.


Fossils

Richard Owen Sir Richard Owen (20 July 1804 – 18 December 1892) was an English biologist, comparative anatomist and paleontologist. Owen is generally considered to have been an outstanding naturalist with a remarkable gift for interpreting fossils. Ow ...
had expertise in comparative anatomy, and his professional judgements revealed a succession of similar species in the same locality, giving Darwin insights which he would later recall as being central to his new views. Owen met Darwin on 29 October 1836 and quickly took on the task of describing these new fossils. At that time the only fully described fossil mammals from South America were three species of ''
Mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...
'' and the gigantic ''Megatherium''. On 9 November, Darwin wrote to his sister that "Some of them are turning out great treasures." The near-complete skeleton from Punta Alta was apparently very closely allied to anteaters, but of the extraordinary size of a small horse. The
rhinoceros A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct specie ...
-sized head bought for two shillings near the city of Mercedes was not a ''megatherium'', but "as far as they can guess, must have been a gnawing animal. Conceive a Rat or a Hare of such a size— What famous Cats they ought to have had in those days!" Over the following years, Owen published descriptions of the most important fossils, naming several as new species. He described the fossils from Punta Alta as including a nearly perfect head and three fragments of heads of '' Megatherium Cuvierii'', the jaw of a related species which Owen named '' Mylodon Darwinii'', and jaws of '' Megalonyx Jeffersonii''. The near-complete skeleton was named '' Scelidotherium'' by Owen, who found that it had most of its bones nearly in their proper relative positions. At the nearby Monte Hermoso beds, numerous rodents included species allied to the
Brazilian tuco-tuco The Brazilian tuco-tuco (''Ctenomys brasiliensis'') is a tuco-tuco species. It is found mainly in the state of Minas Gerais in southeastern Brazil,Fernandes, F. A., R. Fornel, and T. R. O. Freitas. 2012. ''Ctenomys brasiliensis'' Blainville (Ro ...
and the capybara. Owen decided that the fossils of polygonal plates of bony armour found at several locations were not from the ''Megatherium'' as Cuvier's description implied, but from a huge armadillo, as Darwin had briefly thought. Owen found a description of an earlier unnamed specimen which he named '' Glyptodon clavipes'' in 1839. Darwin's find from Punta Alta, a large surface about doubled over with toe bones still inside the folded armour, was identified as a slightly smaller '' Glyptodont'' named '' Hoplophorus'' by Peter Wilhelm Lund in the same year. The huge skull from near Mercedes was named ''
Toxodon ''Toxodon'' (meaning "bow tooth" in reference to the curvature of the teeth) is an extinct genus of South American mammals from the Late Miocene to early Holocene epochs ( Mayoan to Lujanian in the SALMA classification) (about 11.6 million t ...
'' by Owen, and he showed that the "enormous gnawing tooth" from the cliffs of the
Carcarañá River The Carcarañá River (Spanish, Arroyo Saladillo or Río Saladillo) is a river in Argentina. It begins at the confluence of the Río Tercero and the Saladillo River (the lower course of the Río Cuarto) in the south-east of the province of Córd ...
was a molar from this species. The finds near Mercedes also included a large fragment of ''Glyptodont'' armour and a head that Owen initially identified as a '' Glossotherium'', but later decided was a ''Mylodon''. Owen found fragments of the jaw and a tooth of another ''Toxodon'' in the fossils from Punta Alta. The fossils from near Santa Fe included the horse tooth which had puzzled Darwin as it had been previously thought that horses had only come to the Americas in the 16th century, close to a ''
Toxodon ''Toxodon'' (meaning "bow tooth" in reference to the curvature of the teeth) is an extinct genus of South American mammals from the Late Miocene to early Holocene epochs ( Mayoan to Lujanian in the SALMA classification) (about 11.6 million t ...
'' tooth and a tooth of ''Mastodon andium'' (now ''
Cuvieronius ''Cuvieronius'' is an extinct New World genus of gomphothere, named after the French naturalist Georges Cuvier. Alive, specimens typically stood about tall at the shoulder, weighed about and would have superficially resembled a modern elephant ...
hyodon''). Owen confirmed that the horse tooth was of an extinct South American species which he named '' Equus curvidens'', and its age was confirmed by a corroded horse tooth among the Punta Alta fossils. This discovery was later explained as part of the evolution of the horse. The "soft as cheese" ''
Mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...
'' bones at the Paraná River were identified as two gigantic skeletons of ''Mastodon andium'', and mastodon teeth were also identified from Santa Fe and the Carcarañá River. The pieces of spine and a hind leg from Port S. Julian, which Darwin had thought came from "some large animal, I fancy a Mastodon", gave Owen difficulties, as the creature which he named '' Macrauchenia'' appeared to be a "gigantic and most extraordinary pachyderm", allied to the '' Palaeotherium'', but with affinities to the llama and the
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. ...
. The fossils at Punta Alta included a pachyderm tooth which was thought probably came from ''Macrauchenia''.


Footnotes


Notes


References

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External links

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Further reading

*
The Complete Works of Charles Darwin Online The Complete Work of Charles Darwin Online (or Darwin Online) is a freely-accessible website containing the complete print and manuscript works of Charles Darwin, as well as related supplementary material. Overview Darwin Online is a research ...

Darwin Online
Darwin's publications, private papers and bibliography, supplementary works including biographies, obituaries and reviews. Free to use, includes items not in public domain. *; public domain
Darwin Correspondence Project
Text and notes for most of his letters
Darwin in Galapagos: Footsteps to a New World
{{DEFAULTSORT:Second Voyage Of Hms Beagle Voyage on HMS ''Beagle'' History of evolutionary biology Global expeditions 1831 in the United Kingdom 1833 in Argentina 1830s in science HMS Beagle Expeditions from the United Kingdom