Scopoli's shearwater
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Scopoli's shearwater (''Calonectris diomedea'') is a
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
in the petrel family
Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes (or tubenoses), which als ...
. It breeds on rocky islands and on steep coasts in the Mediterranean but outside the breeding season it forages in the Atlantic. It is brownish grey above with darker wings and mostly white below. The bill is pale yellow with a dark patch near the tip. The sexes are alike. It was formerly considered to be
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organis ...
with
Cory's shearwater Cory's shearwater (''Calonectris borealis'') is a large shearwater in the seabird family Procellariidae. It breeds colonially of rocky islands in the eastern Atlantic. Outside the breeding season it ranges widely in the Atlantic. It was formerl ...
.


Taxonomy

Scopoli's shearwater was formally described in 1769 by the Austrian naturalist
Giovanni Antonio Scopoli Giovanni Antonio Scopoli (sometimes Latinized as Johannes Antonius Scopolius) (3 June 1723 – 8 May 1788) was an Italian physician and naturalist. His biographer Otto Guglia named him the "first anational European" and the " Linnaeus of the A ...
. He placed it with the other petrels in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Procellaria Procellaria is a genus of Southern Ocean long-winged seabirds related to prions, and within the order Procellariiformes. The black petrel (''Procellaria parkinsoni'') ranges in the Pacific Ocean, and as far north as Central America. The specta ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''Procellaria diomedea''. Scopoli did not mention a type locality but this was designated in 1946 by the
British Ornithologists' Union The British Ornithologists' Union (BOU) aims to encourage the study of birds ("ornithology") and around the world, in order to understand their biology and to aid their conservation. The BOU was founded in 1858 by Professor Alfred Newton, Henr ...
as the
Tremiti Islands The Isole Tremiti, also called "Isole Diomedee" (Diomedes' Islands, from Greek ''Diomèdee'', Διομήδεες) are an archipelago in the Adriatic Sea, north of the Gargano Peninsula. They constitute a ''"comune"'' of Italy's Province of Foggi ...
in the
Adriatic The Adriatic Sea () is a body of water separating the Italian Peninsula from the Balkan Peninsula. The Adriatic is the northernmost arm of the Mediterranean Sea, extending from the Strait of Otranto (where it connects to the Ionian Sea) to the ...
. Scopoli's shearwater is now placed in the genus ''
Calonectris ''Calonectris'' is a genus of seabirds. The genus name comes from Ancient Greek ''kalos'', "good" and ''nectris'', "swimmer". The genus comprises four large shearwaters. There are two other shearwater genera. ''Puffinus'', which comprises abou ...
'' that was introduced in 1915 by the ornithologists
Gregory Mathews Gregory Macalister Mathews CBE FRSE FZS FLS (10 September 1876 – 27 March 1949) was an Australian-born amateur ornithologist who spent most of his later life in England. Life He was born in Biamble in New South Wales the son of Robert H. M ...
and
Tom Iredale Tom Iredale (24 March 1880 – 12 April 1972) was an English-born ornithologist and malacologist who had a long association with Australia, where he lived for most of his life. He was an autodidact who never went to university and lacked forma ...
. The genus name combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic pe ...
''kalos'' meaning "good" or "noble" with the genus name ''Nectris'' that was used for shearwaters by the German naturalist
Heinrich Kuhl Heinrich Kuhl (17 September 1797 – 14 September 1821) was a German naturalist and zoologist. Kuhl was born in Hanau (Hesse, Germany). Between 1817 and 1820, he was the assistant of professor Th. van Swinderen, docent natural history at the ...
in 1820. The name ''Nectris'' comes from the Ancient Greek ''nēktris'' meaning "swimmer". The specific epithet ''diomedea'' refers to
Diomedes Diomedes (Jones, Daniel; Roach, Peter, James Hartman and Jane Setter, eds. ''Cambridge English Pronouncing Dictionary''. 17th edition. Cambridge UP, 2006.) or Diomede (; grc-gre, Διομήδης, Diomēdēs, "god-like cunning" or "advised by ...
, a hero in
Greek mythology A major branch of classical mythology, Greek mythology is the body of myths originally told by the ancient Greeks, and a genre of Ancient Greek folklore. These stories concern the origin and nature of the world, the lives and activities o ...
. His wife was serially unfaithful while he fought at
Troy Troy ( el, Τροία and Latin: Troia, Hittite: 𒋫𒊒𒄿𒊭 ''Truwiša'') or Ilion ( el, Ίλιον and Latin: Ilium, Hittite: 𒃾𒇻𒊭 ''Wiluša'') was an ancient city located at Hisarlik in present-day Turkey, south-west of Ç ...
, so he left to found a city in Italy. After his death, his distraught friends were turned into white seabirds. The species is considered to be
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
: no
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
are recognised. Scopoli's shearwater and
Cory's shearwater Cory's shearwater (''Calonectris borealis'') is a large shearwater in the seabird family Procellariidae. It breeds colonially of rocky islands in the eastern Atlantic. Outside the breeding season it ranges widely in the Atlantic. It was formerl ...
were previously considered as
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organis ...
. They formed the Cory's shearwater complex (''Calonectris diomedea''). Based on the lack of hybridization and differences in mitochondrial DNA, morphology and vocalization, the complex was split into two separate species. The English name "Cory's shearwater" was transferred to ''Calonectris borealis'' while what was previously the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
became Scopoli's shearwater (''Calonectris diomedea''). Although most ornithological authorities treat Cory's shearwater and Scopoli's shearwater as separate species, the
Cornell Laboratory of Ornithology The Cornell Lab of Ornithology is a member-supported unit of Cornell University in Ithaca, New York, which studies birds and other wildlife. It is housed in the Imogene Powers Johnson Center for Birds and Biodiversity in Sapsucker Woods Sanctuar ...
has chosen not to do so in their updates to ''
The Clements Checklist of Birds of the World ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 202 ...
''.


Description

Scopoli's shearwater is in overall length with a wingspan of . The upperparts are brownish-grey with most feathers fringed with a lighter brown. The wings are a darker brown. The upper tail-coverts are tipped whitish and the tail is dark brown. The underparts are mostly white with a brown border which is most prominent of the trailing edge of the wing. The bill is pale yellow with a dark patch near the tip. The legs and feet are a pale flesh colour. The sexes are similar in appearance but the male is on average slightly larger than the female. See Supplementary Data for morphological measurements. The appearance is very similar to Cory's shearwater and the two species can be difficult to distinguish. The underside of the wing of Scopoli's shearwater has more white on the feathers at the wingtip, in particular the outermost large feather (P10). The
Cape Verde shearwater The Cape Verde shearwater (''Calonectris edwardsii''), or cagarra locally, is a medium-large shearwater, a seabird in the petrel family Procellariidae. It is Endemism in birds, endemic to the Cape Verde archipelago of Macaronesia in the Atlantic ...
is smaller and is significantly darker above.


Distribution and habitat

Scopoli's shearwater breeds on islands in the Mediterranean from the Chafarinas Islands off the Moroccan coast in the west to the
Dodecanese The Dodecanese (, ; el, Δωδεκάνησα, ''Dodekánisa'' , ) are a group of 15 larger plus 150 smaller Greek islands in the southeastern Aegean Sea and Eastern Mediterranean, off the coast of Turkey's Anatolia, of which 26 are inhabited ...
near Turkey in the east. The largest colony is on the rocky island of
Zembra Zembra ( aeb, زمبرة ') is a Tunisian island. The island is a rock formation, and as such contains many 400 metre-high cliffs. It has an area of . Located from El Haouaria and from the port of La Goulette, it is a natural extension of the ...
, off the Tunisian coast. The colony contains between 141,000 and 223,000 breeding pairs which represents more than 75 percent of the global population. Other large colonies are on the island of
Linosa Linosa (; scn, Linusa; ar, نموشة, ''Nammūša'') is one of the Pelagie Islands in the Sicily Channel of the Mediterranean Sea. The island is a part of the Italian comune of Lampedusa e Linosa, part of the province of Agrigento in Sicily, ...
in the
Strait of Sicily The Strait of Sicily (also known as Sicilian Strait, Sicilian Channel, Channel of Sicily, Sicilian Narrows and Pantelleria Channel; it, Canale di Sicilia or the Stretto di Sicilia; scn, Canali di Sicilia or Strittu di Sicilia, ar, مضيق ص ...
, and on the
Balearic Islands The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...
. At the end of October, after the breeding season, Scopoli's shearwaters migrate to the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
and stream out of the Mediterranean through the
Strait of Gibraltar The Strait of Gibraltar ( ar, مضيق جبل طارق, Maḍīq Jabal Ṭāriq; es, Estrecho de Gibraltar, Archaic: Pillars of Hercules), also known as the Straits of Gibraltar, is a narrow strait that connects the Atlantic Ocean to the Medi ...
. They return to the Mediterranean at the end of February. Studies using
light level geolocator A light level geolocator, light-level logger or GLS (global location sensor) is essentially a lightweight, electronic archival tracking device, usually used in bird migration research to map migration routes, identify important staging areas, and s ...
s have found that birds tagged either on the island of Linosa or on the
Pantaleu Es Pantaleu ( Catalan) or El Pantaleu (Spanish) is an uninhabited islet in the Balearic Islands, Spain, located just off the west coast of Majorca. It is currently a natural park, part of the natural park of its neighboring island Dragonera. It ...
islet in the
Balearic Islands The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...
wintered in regions associated with major upwellings in the south east Atlantic. The birds either foraged off the coast of
West Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali ...
in the upwelling associated with the
Canary Current The Canary Current is a wind-driven surface current that is part of the North Atlantic Gyre. This eastern boundary current branches south from the North Atlantic Current and flows southwest about as far as Senegal where it turns west and later j ...
or continued further south and foraged in the Benguela Current off the coast of
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
.


Food and feeding

Scopoli's shearwater mainly feeds on small fish, but it also consumes
cephalopods A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, a ...
and
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
. It feeds by skimming over the surface or by surface feeding but only rarely plunges completely beneath the surface. Sometimes it will follows whales and tuna to pick up food scraps and to catch small fish driven to the surface. It will also scavenge discards from fishing vessels.


References

{{Taxonbar, from=Q216850 Scopoli's shearwater Birds of the Atlantic Ocean Scopoli's shearwater Taxa named by Giovanni Antonio Scopoli