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''Symphyotrichum cordifolium'' (formerly ''Aster cordifolius''), commonly known as common blue wood aster, heartleaf aster, and blue wood-aster, is a
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition ...
flowering plant in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae ...
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and enterta ...
to central and eastern North America. It reaches heights of up to and has bluish daisy-like flowers which bloom late-summer and fall in its range.


Description

''Symphyotrichum cordifolium'' reaches heights of up to . The lower leaves are heart-shaped, while leaves higher on the stem tend to be
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
with more rounded bases. The composite flowers, which have bluish to rarely white
ray floret The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...
s and light yellow
disc floret The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae we ...
s that eventually turn purple, emerge in August and persist into October. File:Symphyotrichum cordifolium 127419951.jpg, ''Symphyotrichum cordifolium'' leaves File:Symphyotrichum cordifolium 99994137.jpg, Flower heads showing involucres File:Symphyotrichum cordifolium 108433505.jpg, Close-up of an involucre showing phyllaries File:Symphyotrichum cordifolium 108433539.jpg, Abaxial side of a leaf File:Symphyotrichum cordifolium 101650600.jpg, Flower heads File:Symphyotrichum cordifolium 117798819.jpg, Showing light blue rays


Chromosomes

''S. cordifolium'' has a
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins ar ...
base number of ''x'' = 8.
Diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respecti ...
and
tetraploid Polyploidy is a condition in which the cells of an organism have more than one pair of (homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contains ...
cytotype A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
s with respective counts of 16 and 32 have been reported.


Taxonomy

''Symphyotrichum cordifolium'' is classified in the
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed betw ...
''Symphyotrichum'',
section Section, Sectioning or Sectioned may refer to: Arts, entertainment and media * Section (music), a complete, but not independent, musical idea * Section (typography), a subdivision, especially of a chapter, in books and documents ** Section sign ...
''Symphyotrichum''. The species was first formally described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
in 1753 as ''Aster cordifolius''.


Hybrids

Where the range of ''Symphyotrichum cordifolium'' overlaps with that of '' S. puniceum'', the
F1 hybrid An F1 hybrid (also known as filial 1 hybrid) is the first filial generation of offspring of distinctly different parental types. F1 hybrids are used in genetics, and in selective breeding, where the term F1 crossbreed may be used. The term is somet ...
named ''Symphyotrichum'' × ''tardiflorum'' can occur. ''Symphyotrichum'' × ''schistosum'' is the hybrid of ''S. cordifolium'' and ''S. laeve'' var. ''laeve''. A hybrid of ''S. cordifolium'' and '' S. shortii'' may occur, and this has been named ''Symphyotrichum'' × ''finkii''. Hybrids with ''S. drummondii'' and ''S. urophyllum'' have been reported but not confirmed.


Distribution and habitat

''Symphyotrichum cordifolium'' occurs from
Manitoba , image_map = Manitoba in Canada 2.svg , map_alt = Map showing Manitoba's location in the centre of Southern Canada , Label_map = yes , coordinates = , capital = Win ...
, east to
Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native En ...
and
Maine Maine () is a U.S. state, state in the New England and Northeastern United States, Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Provinces and territories of Canad ...
, south to
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to t ...
and
Alabama (We dare defend our rights) , anthem = " Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,7 ...
, and west to Oklahoma. It was once introduced in
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost Provinces and territories of Canada, province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include ...
but did not persist. It is an
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived the ...
in
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
and
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...
. It grows primarily in
mesic Mesic may refer to: * Mesic, North Carolina, a town in the United States * Mesic habitat, a type of habitat See also *Mesić (disambiguation) *Mešić Mešić is a Bosnian surname, a patronymic derived from the masculine given name '' Meša'', it ...
sites with soils that are rocky to loamy but generally rich, at heights ranging from sea level along the
coastal plain A coastal plain is flat, low-lying land adjacent to a sea coast. A fall line commonly marks the border between a coastal plain and a piedmont area. Some of the largest coastal plains are in Alaska and the southeastern United States. The Gulf Co ...
up to in the
Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. The ...
. It can be found on open wooded slopes, along the banks of streams, on moist ledges, in swampy woods, along the borders of
beech–maple forest A beech–maple forest or a maple beech forest is a climax mesic closed canopy hardwood forest. It is primarily composed of American beech and sugar maple trees which co-dominate the forest and which are the pinnacle of plant succession in their ...
s and
oak–hickory forest Oak–hickory forest is a type of North American forest ecosystem, and an ecoregion of the Temperate broadleaf and mixed forests Biome. Geography It has a range extending from Rhode Island and southern New York, west to Iowa, and south to Northern ...
s, as well as in clearings, thickets, and along roadsides and ditches. It also can be found in urban areas where it is sometimes considered a weed species.


Ecology


Reproduction

''Symphyotrichum cordifolium'' reproduces
vegetatively Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spec ...
via short rhizomes, as well as via wind-dispersed seeds. The ray florets of species in the ''Symphyotrichum'' genus are exclusively female, each having a pistil but no stamen, while disk florets are bisexual, each with both male and female reproductive parts.


Pests and diseases

Two rusts have been recorded on ''S. cordifolium'': the brown rust ''
Puccinia dioicae ''Puccinia dioicae'' is a plant pathogen that causes rust on goldenrod. It is common in Iceland, where it infects ''Taraxacum'' species and ''Carex capillaris''. Pycniospores and aeciospores are found on ''Taraxacum'' sp., and uredospores and te ...
'' and the red rust ''
Coleosporium asterum is a species of rust fungus in the family Coleosporiaceae. It infects species in the Asteraceae family, such as those in genus ''Aster'' and ''Solidago'', as well as the needle pines ''Pinus contorta'' and '' P. banksiana''. It has been recor ...
''.


Conservation

''Symphyotrichum cordifolium'' has coefficients of conservatism (C-values) in the
Floristic Quality Assessment Floristic Quality Assessment (FQA) is a tool used to assess an area's ecological integrity based on its plant species composition. Floristic Quality Assessment was originally developed in order to assess the likelihood that impacts to an area "woul ...
(FQA) that range from depending on evaluation region. The lower the C-value, the higher tolerance the species has for disturbance and the lesser the likelihood that it is growing in a presettlement natural community. In
the Dakotas The Dakotas is a collective term for the U.S. states of North Dakota and South Dakota. It has been used historically to describe the Dakota Territory, and is still used for the collective heritage, culture, geography, fauna, sociology, econo ...
, for example, ''S. cordifolium'' has a C-value of 8, meaning its populations there are found in high-quality
remnant natural area A remnant natural area, also known as remnant habitat, is an ecological community containing native flora and fauna that has not been significantly disturbed by destructive activities such as agriculture, logging, pollution, development, fire ...
s with little
environmental degradation Environmental degradation is the deterioration of the environment through depletion of resources such as quality of air, water and soil; the destruction of ecosystems; habitat destruction; the extinction of wildlife; and pollution. It is d ...
but can tolerate some periodic disturbance. In contrast, in the Laurentian plains and hills of Maine and
New Brunswick New Brunswick (french: Nouveau-Brunswick, , locally ) is one of the thirteen Provinces and territories of Canada, provinces and territories of Canada. It is one of the three Maritime Canada, Maritime provinces and one of the four Atlantic Canad ...
, it has been given a C-value of 2, meaning its presence in locations of that
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas o ...
provides little confidence of a remnant habitat. ,
NatureServe NatureServe, Inc. is a non-profit organization based in Arlington County, Virginia, US, that provides proprietary wildlife conservation-related data, tools, and services to private and government clients, partner organizations, and the public. Nat ...
listed ''Symphyotrichum cordifolium'' as Secure (G5) worldwide, Critically Imperiled (S1) in
Kansas Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to ...
, and Vulnerable (S3) in
Delaware Delaware ( ) is a state in the Mid-Atlantic region of the United States, bordering Maryland to its south and west; Pennsylvania to its north; and New Jersey and the Atlantic Ocean to its east. The state takes its name from the adjacen ...
and
Missouri Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee ...
.


Uses


Medicinal

The
Ojibwe The Ojibwe, Ojibwa, Chippewa, or Saulteaux are an Anishinaabe people in what is currently southern Canada, the northern Midwestern United States, and Northern Plains. According to the U.S. census, in the United States Ojibwe people are one of ...
have used ''S. cordifolium'' to make an incense to attract deer.


Gardening

''Symphyotrichum cordifolium'' is cultivated as a garden plant under its current name and the older name ''Aster cordifolius''. Several cultivars have been selected for garden use, and the following have achieved the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (No ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
: *'Chieftain' has "light mauve-blue ray florets." *'Little Carlow' (''S. cordifolium'' hybrid) with "abundant violet-blue" rays. *‘Photograph’ (''S. cordifolium'' hybrid) has "pale lilac-blue flowers." *'Sweet Lavender' with "lavender-blue flowers."


Citations


References

* * * * * * * * * * * * * * * * * * {{Taxonbar, from1=Q4050493, from2=Q21871600 cordifolium Flora of Canada Flora of the United States Plants used in traditional Native American medicine Plants described in 1753 Taxa named by Carl Linnaeus