Styphelia Williamsiorum

''Styphelia'' is a genus of shrubs in the family Ericaceae, native from Mainland Southeast Asia, Indo-China through the Pacific to Australia. Most have minute or small leaves with a sharp tip, single, tube-shaped flowers arranged in leaf wikt:axil, axils and with the ends of the petals rolled back with hairs in the inside of the tube.

Description

Plants in the genus ''Styphelia'' are usually erect or spreading shrubs that have egg-shaped, elliptical or oblong, more or less Sessility (botany), sessile leaves with many fine, almost parallel veins and a sharp point on the tip. The flowers are usually arranged singly in leaf axils with small bracts grading to larger Bract#Bracteole, bracteoles at the base and five, usually coloured sepals. The petals are fused to form a cylindrical tube with their tips rolled back. The inside of the petal tube is hairy and the five stamens and thread-like Style (botany), style extend beyond the end of the tube. The fruit is a drupe with a dry or slightly fleshy Fruit anatomy#Mesocarp, mesocarp and a hard Fruit anatomy#Endocarp, endocarp.

Taxonomy and naming

The genus ''Styphelia'' was first formally described in 1793 by James Edward Smith (botanist), James Edward Smith in his book ''A Specimen of the Botany of New Holland''. The name ''Styphelia'' is from an ancient Greek word meaning "tough" or "harsh", referring to the "stiff, prickly leaves and general habit" of the genus.

Species list

The following is a list of species of ''Styphelia'' accepted by Plants of the World Online as at October 2024:

* ''Styphelia abnormis'' (Sond.) F.Muell. (Northern Borneo to New Guinea)
* ''Styphelia abscondita'' J.J.Sm. (New Guinea)
* ''Styphelia acervata'' (Michael Clyde Hislop, Hislop & Annette Jane Gratton Wilson, A.J.G.Wilson) Hislop, Darren M. Crayn, Crayn & Caroline Puente-Lelievre, Puente-Lel. (W.A.)
* ''Styphelia acuminata'' (R.Br.) Spreng. (N.T.)
* ''Styphelia adscendens'' R.Br. - golden heath (N.S.W., S.A., Vic., Tas.)
* ''Styphelia allittii'' (F.Muell.) F.Muell. (W.A.)
* ''Styphelia angustiflora'' Hislop & Puente-Lel. (W.A.)
* ''Styphelia angustifolia'' DC. (N.S.W.)
* ''Styphelia annulata'' Hislop (W.A.)
* ''Styphelia anomala'' Hislop (W.A.)
* ''Styphelia appressa'' (R.Br.) Spreng. (N.S.W., Qld., Vic.)
* ''Styphelia attenuata'' (A.Cunn.) F.Muell. – grey beard-heath (N.S.W., Qld., Vic.)
* ''Styphelia balansae'' Robert Virot, Virot (New Caledonia)
* ''Styphelia biflora'' (R.Br.) Spreng. (N.S.W., Qld., Vic.)
* ''Styphelia blackallii'' Hislop (W.A.)
* ''Styphelia blakei'' Hislop, Crayn & Puente-Lel. (Qld.)
* ''Styphelia blepharolepis'' F.Muell. (W.A.)
* ''Styphelia brachygyna'' Hislop (W.A.)
* ''Styphelia bracteolosa'' Hislop (W.A.)
* ''Styphelia brevicuspis'' (Benth.) F.Muell. (W.A.)
* ''Styphelia breviflora'' (F.Muell.) F.Muell. (W.A.)
* ''Styphelia browniae'' Hislop (W.A.)
* ''Styphelia capillaris'' Hislop & Puente-Lel. – Horts' styphelia (W.A.)
* ''Styphelia carolineae'' Hislop (W.A.)
* ''Styphelia caudata'' Hislop (W.A.)
* ''Styphelia cernua'' Hislop & Puente-Lel. (W.A.)
* ''Styphelia chlorantha'' Hislop & A.J.G.Wilson (W.A.)
* ''Styphelia ciliosa'' Hislop & Puente-Lel. (W.A.)
* ''Styphelia clelandii'' (Edwin Cheel, Cheel) J.H.Willis – Cleland's bearded-heath (S.A., Vic.)
* ''Styphelia coelophylla'' (A.Cunn. ex DC.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia cognata'' A.R.Bean (Qld.)
* ''Styphelia compacta'' (R.Br.) Spreng. (W.A.)
* ''Styphelia conchifolia'' (Arne Strid, Strid) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia concinna'' (Benth.) F.Muell. (W.A.)
* ''Styphelia conferta'' (Benth.) F.Muell. – Torrington beard-heath (N.S.W., Qld.)
* ''Styphelia conostephioides'' (DC.) F.Muell. (W.A.)
* ''Styphelia cordifolia'' (Lindl.) F.Muell. – heart-leaved beard-heath (N.S.W., S.A., Vic., W.A.)
* ''Styphelia corynocarpa'' (Sond.) F.Muell. (W.A.)
* ''Styphelia coryphila'' (André Guillaumin, Guillaumin) Sleumer (New Caledonia)
* ''Styphelia crassiflora'' F.Muell. (W.A.)
* ''Styphelia crassifolia'' (Sond.) F.Muell. (W.A.)
* ''Styphelia cuspidata'' (R.Br.) Spreng. (Qld.)
* ''Styphelia cymbiformis'' (A.Cunn. ex DC.) F.Muell. (W.A.)
* ''Styphelia cymbulae'' (Labill.) Spreng. (Fiji, New Caledonia, Vanuatu)
* ''Styphelia cylindrica'' Hislop (W.A.)
* ''Styphelia dammarifolia'' (Brongn. & Jean Antoine Arthur Gris, Gris) F.Muell. (New Caledonia)
* ''Styphelia decussata'' Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia deformis'' (R.B.) Spreng. (N.S.W., Qld.)
* ''Styphelia densifolia'' Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia deserticola'' Hislop (W.A.)
* ''Styphelia dielsiana'' (E.Pritz.) Sleumer (W.A.)
* ''Styphelia discolor'' (Sond.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia disjuncta'' Hislop & Puente-Lel. (W.A.)
* ''Styphelia echinulata'' Hislop (W.A.)
* ''Styphelia enervia'' (Guillaumin) Sleumer (New Caledonia)
* ''Styphelia epacridis'' (DC.) F.Muell. (W.A.)
* ''Styphelia erectifolia'' Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia ericoides'' Sm. – pink beard-heath (N.S.W., Qld., S.A., Tas., Vic.)
* ''Styphelia erubescens'' F.Muell. (W.A.)
* ''Styphelia esquamata'' (R.Br.) Spreng. – swamp beard-heath (N.S.W., Qld., Tas., Vic.)
* ''Styphelia exarata'' Hislop (W.A.)
* ''Styphelia exarrhena'' (F.Muell.) F.Muell. - desert styphelia (S.A., Vic.)
* ''Styphelia exilis'' Hislop (W.A.)
* ''Styphelia exolasia'' F.Muell. – Woronora beard-heath (N.S.W.)
* ''Styphelia exserta'' (F.Muell.) Sleumer (W.A.)
* ''Styphelia filamentosa'' Hislop & Puente-Lel. (W.A.)
* ''Styphelia filifolia'' Hislop & Puente-Lel. (W.A.)
* ''Styphelia flavescens'' (Sond.) F.Muell. (W.A.)
* ''Styphelia fletcheri'' (Joseph Maiden, Maiden & Ernst Betche, Betche) Maiden & Betche (N.S.W., Qld., Vic.)
* ''Styphelia flexifolia'' (R.Br.) Spreng. (Qld.)
* ''Styphelia foliosa'' (Sond.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia forbesii'' Sleumer (Lesser Sunda Islands)
* ''Styphelia geniculata'' Crayn (Qld.)
* ''Styphelia glaucifolia'' (W.Fitzg.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia globosa'' Hislop (W.A.)
* ''Styphelia grandiflora'' (Leslie Pedley, Pedley) Hislop, Crayn & Puente-Lel. (Qld.)
* ''Styphelia graniticola'' Hislop (W.A.)
* ''Styphelia hainesii'' F.Muell. (W.A.)
* ''Styphelia hamulosa'' (E.Pritz.) Sleumer (W.A.)
* ''Styphelia hispida'' (E.Pritz.) Sleumer (W.A.)
* ''Styphelia howatharra'' Hislop (W.A.)
* ''Styphelia humifusa'' (Cav.) Pers. – native cranberry, cranberry heath (N.S.W, S.A., Tas., Vic.)
* ''Styphelia hyalina'' Hislop (W.A.)
* ''Styphelia imbricata'' (R.Br.) Spreng. (Qld.)
* ''Styphelia incerta'' Hislop (W.A.)
* ''Styphelia inframediana'' Hislop (W.A.)
* ''Styphelia inopinata'' (Hislop) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia insularis'' (A.Cunn. ex DC.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia intertexta'' A.S.George (W.A.)
* ''Styphelia intricata'' Hislop (W.A.)
* ''Styphelia javanica'' (Zoll. & Moritzi) J.J.Sm. (Java)
* ''Styphelia kalbarriensis'' Hislop & E.A.Br. (W.A.)
* ''Styphelia kingiana'' F.Muell. (W.A.)
* ''Styphelia laeta'' R.Br. (N.S.W.) – five corners
* ''Styphelia lanata'' Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia lavarackii'' (Pedley) Hislop, Crayn & Puente-Lel. (Qld.)
* ''Styphelia leptantha'' (Benth.) F.Muell. (W.A.)
* ''Styphelia leptospermoides'' (R.Br.) Spreng. (N.S.W., Qld.)
* ''Styphelia lissanthoides'' (F.Muell.) F.Muell. (S.A., W.A.)
* ''Styphelia longifolia'' R.Br. – long-leaf styphelia (N.S.W.)
* ''Styphelia longissima'' Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia longistylis'' (Brongn. & Gris) Sleumer (New Caledonia)
* ''Styphelia lucens'' A.R.Bean (Qld.)
* ''Styphelia macrocalyx'' (Sond.) F.Muell. – Swan berry (W.A.)
* ''Styphelia macrocarpa'' (Schltr.) Sleumer (New Caledonia)
* ''Styphelia madida'' Hislop (W.A.)
* ''Styphelia malayana'' (William Jack (botanist), Jack) J.J.Sm. (Borneo, Cambodia, Malaya, Myanmar, New Guinea, Sumatra, Thailand, Vietnam)
* ''Styphelia margarodes'' (R.Br.) Spreng. (N.S.W., Qld.)
* ''Styphelia marginata'' (W.Fitzg.) Hislop, Crayn & Puente-Lel. – thick-margined leucopogon (W.A.)
* ''Styphelia melaleucoides'' F.Muell. (W.A.)
* ''Styphelia microcalyx'' (Sond.) F.Muell. native cranberry (W.A.)
* ''Styphelia microdonta'' (F.Muell. ex Benth) F.Muell. (W.A.)
* ''Styphelia mitchellii'' (Benth.) F.Muell. (Qld.)
* ''Styphelia multiflora'' Spreng. (Chatham Island, N.S.W., Vic., W.A.)
* ''Styphelia mutica'' (R.Br.) F.Muell. – blunt beard-heath (N.S.W., Qld.)
* ''Styphelia nana'' (Murray Ian Dawson, M.I.Dawson & Peter Brian Heenan, Heenan) Hislop, Crayn & Puente-Lel. (South Island, New Zealand South Island)
* ''Styphelia neoanglica'' (F.Muell. ex Benth.) F.Muell. – New England beard heath (N.S.W., Qld.)
* ''Styphelia nesophila'' (DC.) Sleumer – sharp beard-heath, pātōtara, dwarf mingimingi (N.S.W., New Zealand, Tasmania, Victoria)
* ''Styphelia nitens'' Sleumer (W.A.)
* ''Styphelia oblongifolia'' (A.J.G.Wilson & Hislop) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia obtecta'' (Benth.) F.Muell. (W.A.)
* ''Styphelia pallens'' Hislop (W.A.)
* ''Styphelia pallida'' (R.Br.) Spreng. – kick bush (W.A.)
* ''Styphelia pancheri'' (Brongn. & Gris) F.Muell. (New Caledonia)
* ''Styphelia papillosa'' Hislop (W.A.)
* ''Styphelia pendula'' (R.Br.) Spreng. (W.A.)
* ''Styphelia pentapogona'' F.Muell. (W.A.)
* ''Styphelia perileuca'' Jocelyn Marie Powell, J.M.Powell - montane green five-corners (N.S.W.)
* ''Styphelia piliflora'' Crayn (Qld.)
* ''Styphelia planiconvexa'' Hislop (W.A.)
* ''Styphelia planifolia'' (Sond.) Sleumer (W.A.)
* ''Styphelia platyneura'' Hislop (W.A.)
* ''Styphelia pogonocalyx'' (F.Muell. ex Benth.) F.Muell. (W.A.)
* ''Styphelia propinqua'' (R.Br.) Spreng. (W.A.)
* ''Styphelia prostrata'' (R.Br.) F.Muell. (W.A.)
* ''Styphelia psiloclada'' J.M.Powell (N.S.W.)
* ''Styphelia psilopus'' (Stschegl.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia pubescens'' (S.Moore) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia quartzitica'' Hislop (W.A.)
* ''Styphelia racemulosa'' (DC.) F.Muell. (W.A.)
* ''Styphelia rectiloba'' Hislop (W.A.)
* ''Styphelia recurva'' Hislop (W.A.)
* ''Styphelia recurvisepala'' (C.T.White) Sleumer (N.S.W., Qld.)
* ''Styphelia retrorsa'' Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia rigida'' (A.Cunn. ex DC.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia riparia'' (N.A.Wakef.) J.H.Willis (Vic.)
* ''Styphelia roseola'' Hislop (W.A.)
* ''Styphelia rotundifolia'' (R.Br.) Spreng. (W.A.)
* ''Styphelia rufa'' (Lindl.) F.Muell. – spoon-leaf beard-heath, ruddy bearded-heath (N.S.W., S.A., Vic.)
* ''Styphelia rupicola'' (C.T.White) Sleumer (Qld.)
* ''Styphelia ruscifolia'' (R.Br.) Spreng. (N.T., Qld.)
* ''Styphelia saxicola'' Hislop (W.A.)
* ''Styphelia scabrella'' Hislop (W.A.)
* ''Styphelia serratifolia'' (DC.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia setigera'' (R.Br.) Spreng. (N.S.W., Vic.)
* ''Styphelia sieberi'' (DC.) Hislop, Crayn & Puente-Lel. – prickly beard-heath (N.S.W., Qld., Vic.)
* ''Styphelia sonderensis'' (J.H.Willis) Hislop, Crayn & Puente-Lel. – Mount Sonder beard-heath (N.T.)
* ''Styphelia stomarrhena'' (Sond.) Sleumer – red swamp cranberry (W.A.)
* ''Styphelia striata'' (R.Br.) Spreng. (W.A.)
* ''Styphelia strongylophylla'' (F.Muell.) F.Muell. (W.A.)
* ''Styphelia subglauca'' Hislop (W.A.)
* ''Styphelia subulata'' (F.Muell.) Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia sulcata'' Hislop & Puente-Lel. (W.A.)
* ''Styphelia tamminensis'' (E.Pritz.) Sleumer (W.A.)
* ''Styphelia tecta'' (R.Br.) Spreng. (W.A)
* ''Styphelia tenuiflora'' Lindl. - common pinheath (W.A.)
* ''Styphelia tortifolia'' Hislop, Crayn & Puente-Lel. (W.A.)
* ''Styphelia trichostyla'' (J.M.Powell) Hislop, Crayn & Puente-Lel. – daphne heath (N.S.W., Qld.)
* ''Styphelia triflora'' Henry Cranke Andrews, Andrews - pink five-corners (Qld., N.S.W., A.C.T.)
* ''Styphelia tubiflora'' Sm. - red five-corners (N.S.W.)
* ''Styphelia undulata'' Hislop (W.A.)
* ''Styphelia veillonii'' Virot (New Caledonia)
* ''Styphelia viridis'' Andrews – green five corners (N.S.W., Qld.)
* ''Styphelia wetarensis'' J.J.Sm. (Lesser Sunda Island)
* ''Styphelia williamsiorum'' Hislop & Puente-Lel. (W.A.)
* ''Styphelia woodsii'' (F.Muell.) F.Muell. – nodding beard-heath (S.A., Vic., W.A.)
* ''Styphelia xerophylla'' (DC.) F.Muell. (W.A.)
* ''Styphelia yorkensis'' (Pedley) Hislop, Crayn & Puente-Lel. (Qld.)

Distribution

Species of ''Styphelia'' occur in all Australian mainland states, the Northern Territory and the Australian Capital Territory. Other species are found northwards to Mainland Southeast Asia, Indo-China.

References

{{Taxonbar, from=Q73917

Styphelia,
Ericaceae genera
Ericales of Australia