''Stratesaurus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of small-bodied rhomaleosaurid

plesiosaur The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...

known from the

Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...

period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...

(most likely earliest

Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic Period of the geologic timescale. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). The Hettangian follows the Rhaetian (part of the Triass ...

stage) of the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...

. It contains a single species, ''S. taylori''. It was a small plesiosaur, with a skull length of and a body length of .

Discovery

''Stratesaurus'' is known from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

specimen OUMNH J.10337, a dorsoventrally crushed but nearly complete

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

, and three-dimensionally preserved partial

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...

including anterior cervical and pectoral vertebrae, a partial

hindlimb A hindlimb or back limb is one of the paired articulated appendages ( limbs) attached on the caudal ( posterior) end of a terrestrial tetrapod vertebrate's torso.http://www.merriam-webster.com/medical/hind%20limb, Merriam Webster Dictionary-H ...

and ilium. The specimen was acquired by the OUMNH in 1874 from the collection of Thomas Hawkins.

GSM The Global System for Mobile Communications (GSM) is a family of standards to describe the protocols for second-generation (2G) digital cellular networks, as used by mobile devices such as mobile phones and Mobile broadband modem, mobile broadba ...

26035 was referred to ''S. taylori'' because it shares one autapomorphy and other characters with the holotype specimen. It consists of a skull and some anterior cervical vertebrae. AGT 11 was also referred to ''S. taylori''. Although it does nor show the autapomorphies of ''S. taylori'', it is indistinguishable from both OUMNH J.10337 and GSM 26035, and it is possible to distinguish it from similar taxa, like ''

Avalonnectes ''Avalonnectes'' is an extinct genus of small-bodied rhomaleosaurid known from the Early Jurassic period (most likely earliest Hettangian stage) of the United Kingdom. It contains a single species, ''A. arturi''. Discovery ''Avalonnectes'' is k ...

''. All specimens were collected at

Street A street is a public thoroughfare in a city, town or village, typically lined with Building, buildings on one or both sides. Streets often include pavements (sidewalks), pedestrian crossings, and sometimes amenities like Street light, streetligh ...

, of

Somerset Somerset ( , ), Archaism, archaically Somersetshire ( , , ) is a Ceremonial counties of England, ceremonial county in South West England. It is bordered by the Bristol Channel, Gloucestershire, and Bristol to the north, Wiltshire to the east ...

, from the Pre-''Planorbis'' beds of the

Blue Lias Formation The Blue Lias is a geological formation in southern, eastern and western England and parts of South Wales, part of the Lias Group. The Blue Lias consists of a sequence of limestone and shale layers, laid down in latest Triassic and early Jurassi ...

of the

Lower Lias Group The Lias Group or Lias is a lithostratigraphy, lithostratigraphic unit (a sequence of rock strata) found in a large area of western Europe, including the British Isles, the North Sea, the Low Countries and the north of Germany. It consists of Sed ...

. These beds likely occur below the first occurrence of the ammonite '' Psiloceras planorbis''. Thus, they probably fall within the earliest Hettangian ''P. tilmanni'' Chronozone, which is about 199.6-198 million years old, immediately following the

Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...

–

Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...

Boundary. Plesiosaurs fossils which were discovered at Street represent the earliest known occurrence of the

Plesiosauria The Plesiosauria or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period, possibly in the Rhaetian stage, about 203 million year ...

. Hence, ''Stratesaurus'' is one of the oldest plesiosaurs to date.

Description

''Stratesaurus'' is a small-bodied rhomaleosaurid, with a skull length of in the holotype. Its snout is not constricted, and it has five of tooth sockets in its

premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammals h ...

and sixteen in the

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

. In the front part of the neck, there are strongly developed backwards-pointing projections on the prezygapophyses, an

autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a Synapomorphy, derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or Outgroup (cladistics), outgroup taxa, not ...

(unique derived feature) of the genus. Also autapomorphic are the short lengths of the centra (

vertebral bodies Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...

) in the shoulder region.

Phylogeny

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

analysis performed by Benson ''et al.'' (2012) found it to be the

basalmost In phylogenetics, basal is the direction of the ''base'' (or root) of a rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to th ...

rhomaleosaurid. The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below shows ''Stratesaurus'' phylogenetic position among other plesiosaurs following Benson ''et al.'' (2012).

Etymology

''Stratesaurus'' was first described and named by

Roger B. J. Benson Roger is a masculine given name, and a surname. The given name is derived from the Old French personal names ' and '. These names are of Germanic languages">Germanic origin, derived from the elements ', ''χrōþi'' ("fame", "renown", "honour") ...

, Mark Evans and Patrick S. Druckenmiller in

2012 2012 was designated as: *International Year of Cooperatives *International Year of Sustainable Energy for All Events January *January 4 – The Cicada 3301 internet hunt begins. * January 12 – Peaceful protests begin in the R ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Stratesaurus taylori''. The generic name is derived from "Strate", the name for

as it recorded in the

Domesday Book Domesday Book ( ; the Middle English spelling of "Doomsday Book") is a manuscript record of the Great Survey of much of England and parts of Wales completed in 1086 at the behest of William the Conqueror. The manuscript was originally known by ...

and from

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

''sauros'', meaning "lizard". The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

honors the paleontologist Michael A. Taylor, who performed acid preparation of the holotype.

References

{{Taxonbar, from=Q5492579 Prehistoric life of Europe Early Jurassic plesiosaurs of Europe Fossil taxa described in 2012 Rhomaleosauridae Sauropterygian genera

Discovery

Description

Phylogeny

Etymology

See also

References