
A squamulose lichen is a
lichen that is composed of small, often overlapping "scales" called
squamules.
If they are raised from the
substrate
Substrate may refer to:
Physical layers
*Substrate (biology), the natural environment in which an organism lives, or the surface or medium on which an organism grows or is attached
** Substrate (locomotion), the surface over which an organism lo ...
and appear leafy, the lichen may appear to be a
foliose lichen, but the underside does not have a "skin" (
cortex), as foliose lichens do.
Squamulose lichens are composed of flattish units that are usually tightly clustered. They are like an intermediate between
crustose and
foliose lichens.

Examples of squamulose lichen include ''
Vahliella leucophaea'', ''
Cladonia subcervicornis'' and ''
Lichenomphalia hudsoniana''.
References
Lichenology
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