The spotted sandpiper (''Actitis macularius'') is a small shorebird. Together with its

sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

the common sandpiper (''A. hypoleucos''), it makes up the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

'' Actitis''. They replace each other geographically; stray birds may settle down with breeders of the other species and hybridize.

Taxonomy

The spotted sandpiper was formally described by the Swedish naturalist

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

in 1766 in the twelfth edition of his ''

Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...

'' under the

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...

''Tringa macularia''. The type locality is

Pennsylvania Pennsylvania, officially the Commonwealth of Pennsylvania, is a U.S. state, state spanning the Mid-Atlantic (United States), Mid-Atlantic, Northeastern United States, Northeastern, Appalachian, and Great Lakes region, Great Lakes regions o ...

. The species is now placed together with common sandpiper in the

'' Actitis'' and was introduced in 1811 by the German zoologist Johann Illiger. The genus name ''Actitis'' is from the

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

''aktites'' meaning "coast-dweller" from ''akte'' meaning "coast". The specific epithet ''macularius'' is

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

meaning "spotted". The species is

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

: no

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

are recognised.

Description

Spotted Sandpiper - 2017-08-17 - Andy Mabbett - 02

Adults have short yellowish legs and an orange bill with a dark tip. The body is brown on top and white underneath with black spots. These spots vary in degree over the course of spotted sandpipers' lives, becoming especially prevalent around the breeding season. The overall health of spotted sandpipers may be suggested by the "spottiness" of an individual. Generally, females with more "spottiness" were healthier than those who did not have as many spots. The condition of males based on the amount of spots they exhibit is yet to be determined. Additionally, spot size gets smaller and the spot shape becomes more irregularly shaped as age increases. Spotted sandpipers also feature a white supercilium. Non-breeding

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s, depicted below, do not have the spotted underparts, and are very similar to the common sandpiper of

Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...

; the main difference is the more washed-out wing pattern visible in flight and the normally light yellow legs and feet of the spotted sandpiper. The ''Actitis'' species have a distinctive stiff-winged flight low over the water. They also have a distinctive walk in which their tails bob up and down. Males and females exhibit similar physical measurements, but differ in weight; females tend to be about 20-25% heavier than males. Measurements: * Length: 7.1-7.9 in (18–20 cm) * Weight: 1.2-1.8 oz (34-50 g) * Wingspan: 14.6-15.8 in (37–40 cm)

Distribution

Spotted sandpipers are a philopatric species. Their breeding habitat is near fresh water across most of

Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...

and the

United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...

. They migrate to the southern United States, the

Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...

, and

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

, and are very rare vagrants to western

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

. These are not gregarious birds and are seldom seen in flocks. Spotted sandpipers are the most widespread species of their kind in North America due to their high breeding rates and their ability to adapt to various environmental pressures.

Behaviour

Breeding

Breeding grounds are chosen based on various environmental factors but tend to be in the proximity of bodies of water that offer some coverage from vegetation. Successful breeding sites may be used repeatedly until either the site becomes physically unsuitable (from overgrown vegetation or flooding) or predation becomes too severe. The females usually arrive at breeding grounds before the males do and establish their territories. Males arrive at breeding sites later but it is uncertain whether they will arrive at the same breeding sites that some females have chosen. The search for mates amongst female spotted sandpipers is much more competitive than finding potential mates for males. During each summer breeding season, females may mate with and lay clutches for more than one male, leaving incubation to them. This is called

polyandry Polyandry (; ) is a form of polygamy in which a woman takes two or more husbands at the same time. Polyandry is contrasted with polygyny, involving one male and two or more females. If a marriage involves a plural number of "husbands and wives ...

. Male spotted sandpipers take care of the offspring, both before and after the offspring hatch. They will incubate their eggs for about 20–23 days. Male parents of first clutches may father chicks in later male's clutches, probably due to sperm storage within female reproductive tracts, which is common in birds. Females that fail to find additional mates usually help incubate and rear chicks. "Prior to incubation, blood plasma concentrations of testosterone and dihydrotestosterone are substantially higher in males than in females" and these levels plummet 25-fold in males as incubation proceeds.Nelson RJ. 2005. ''Introduction to Behavioral Endocrinology''. Sinauer Associates: Massachusetts. p 115. Additionally, mated females have testosterone concentrations that are 7 times higher than those of unmated females. Due to their polyandrous behavior, spotted sandpipers tend to produce more offspring compared to other species of sandpipers.

Food and feeding

These birds forage on the ground or in water, picking up food by sight. They may also catch insects in flight. They eat insects (such as

flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwin ...

beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...

grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are amongst what are possibly the most ancient living groups of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grassh ...

mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the order ...

midge A midge is any small fly, including species in several family (biology), families of non-mosquito nematoceran Diptera. Midges are found (seasonally or otherwise) on practically every land area outside permanently arid deserts and the frigid ...

cricket Cricket is a Bat-and-ball games, bat-and-ball game played between two Sports team, teams of eleven players on a cricket field, field, at the centre of which is a cricket pitch, pitch with a wicket at each end, each comprising two Bail (cr ...

s and

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...

s), crustaceans and other

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

s (such as

spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...

snail A snail is a shelled gastropod. The name is most often applied to land snails, terrestrial molluscs, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gas ...

s, other

mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...

s, and

worm Worms are many different distantly related bilateria, bilateral animals that typically have a long cylindrical tube-like body, no limb (anatomy), limbs, and usually no eyes. Worms vary in size from microscopic to over in length for marine ...

s), as well as small

fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...

and

carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...

. As they forage, they can be recognized by their constant nodding and teetering.

Predators

The main predators of spotted sandpipers include raptors, mustelids, mice, and gulls. Most of these predators target the sandpipers' chicks and eggs.

Conservation status

Although there has been some decline in the population of spotted sandpipers, their conservation status is currently of least concern. However, the decrease in the population of spotted sandpipers is not projected to slow or stop in the future. The destruction of their natural habitats due to increasing wildfires causes problems for breeding and raising offspring. Additionally, the gradual increase in temperatures poses a problem for newborn sandpipers.

Etymology

The word Actitis comes from the Greek word for "coast-dweller", while the word macularius comes from the Latin word for "spotted".

Gallery

File:Actitis-macularia-004.jpg, Front view File:Spotted Sandpiper (non-breeding plumage) (32877802088).jpg, Non-breeding plumage File:Brodziec_plamisty_1.jpg, Spotted sandpiper hunting in the Wallkill River Wildlife Refuge in

New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...

and

New York (state) New York, also called New York State, is a U.S. state, state in the northeastern United States. Bordered by New England to the east, Canada to the north, and Pennsylvania and New Jersey to the south, its territory extends into both the Atlanti ...

File:Spotted Sandpiper (8705161641).jpg, Spotted sandpiper foraging in Fox River Grove,

Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. It borders on Lake Michigan to its northeast, the Mississippi River to its west, and the Wabash River, Wabash and Ohio River, Ohio rivers to its ...

Video

References

External links

Spotted Sandpiper Species Account
- Cornell Lab of Ornithology

- USGS Patuxent Bird Identification InfoCenter * * * {{Taxonbar, from=Q389691

spotted sandpiper The spotted sandpiper (''Actitis macularius'') is a small shorebird. Together with its sister species the common sandpiper (''A. hypoleucos''), it makes up the genus ''Actitis''. They replace each other geographically; stray birds may Hybridisati ...

Birds of North America Birds of the Caribbean Birds of the Dominican Republic Birds of Central America Wintering birds of South America