Solanum Cheesmaniae
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''Solanum cheesmaniae'' is one of two main species of wild
tomato The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible Berry (botany), berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originate ...
es found on the
Galápagos Islands The Galápagos Islands () are an archipelago of volcanic islands in the Eastern Pacific, located around the equator, west of the mainland of South America. They form the Galápagos Province of the Republic of Ecuador, with a population of sli ...
. This species is the one most commonly called the Galapagos tomato. It is a wild tomato that evolved on the famous Galapagos Islands, the place where
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
noted the structural difference between local
finch The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches generally have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where the ...
es,
iguana ''Iguana'' (, ) is a genus of herbivorous lizards that are native to tropical areas of Mexico, Central America, South America, and the Caribbean. The genus was first described by Austrian naturalist Josephus Nicolaus Laurenti, J.N. Laurenti in ...
s, and
barnacle Barnacles are arthropods of the subclass (taxonomy), subclass Cirripedia in the subphylum Crustacean, Crustacea. They are related to crabs and lobsters, with similar Nauplius (larva), nauplius larvae. Barnacles are exclusively marine invertebra ...
s, leading him to identify
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the Heredity, heritable traits characteristic of a population over generation ...
as a possible source of the
origin of species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life'')The book's full original title was ''On the Origin of Species by M ...
; he prepared a
herbarium A herbarium (plural: herbaria) is a collection of preserved plant biological specimen, specimens and associated data used for scientific study. The specimens may be whole plants or plant parts; these will usually be in dried form mounted on a sh ...
sheet of the species on his visit. It often gets confused with the other native species, ''
Solanum galapagense ''Solanum galapagense'' is a wild tomato in the family Solanaceae section ''Lycopersicon'' and is one of two tomato species endemic to the Galápagos Islands, 500 miles west of Ecuador. It was at these islands where Charles Darwin noted the struc ...
'', which is similar but slightly different having more bushy smelly foliage and having smaller hairier orange fruits. It also gets confused with natural omestic tomato x ''S. cheesmaniae''hybrids and 'Solanum pimpinellifolium'' x (''S. cheesmaniae'' or ''S. galapagense'')hybrids because of introduction of domestic tomatoes to the Galapagos Islands, and the presence of ''Solanum pimpinellifolium'' on the islands and natural hybridization occurring between them.Local extirpation is pervasive among historical populations of Galápagos endemic tomatoes
/ref> Although listed as a
Least-concern species A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
by the IUCN, recent observations suggest the species may in fact be threatened. This tomato is smaller and more pale than the oldest mainland cultivars, both in its fruit and leaves, but its flavor is similar. It can also be crossbred with it, which has been done professionally to impart its unusual disease resistances to some commercial cultivars.


Description

''Solanum cheesmaniae'' is a robust,
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
,
herbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition o ...
that at first grows erect, later lying. It reaches up to high; its foliage reaches a similar
diameter In geometry, a diameter of a circle is any straight line segment that passes through the centre of the circle and whose endpoints lie on the circle. It can also be defined as the longest Chord (geometry), chord of the circle. Both definitions a ...
. It is often found in rocky coastal sites. The grayish stems become woody at the base and reach a diameter of . The dense, velvety coat consists of white, single row, non-glandular
trichomes Trichomes (; ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...
up to 0.5 mm in length and occasional short-row, glandular trichomes with four or eight digits. There are varying breeds of galapagos tomato, which can be differentiated by one to four-pinnate leaves, and habits of growing on the shore or inland.Relationships, origin, and diversity of Galápagos tomatoes: implications for the conservation of natural populations
/ref>


Distribution and ecology

The species occurs on the islands of Galapagos, 500 miles west of
Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...
. They grow in extremely wet, rocky areas from sea level to up to several hundred feet. The flowers and fruits appear consistently throughout the year, but there is a noticeable increase in flowering between September and October. ''Solanum cheesmaniae'' is thought to have diverged from its closest relatives, ''
Solanum lycopersicum The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originated from and was d ...
'' and ''
Solanum pimpinellifolium ''Solanum pimpinellifolium'', commonly known as the currant tomato or pimp, is a wild species of tomato native to Ecuador and Peru but naturalized elsewhere, such as the Galápagos Islands. Its small fruits are edible, and it is commonly grown in ...
'', less than 500,000 years ago.Phylogenomics Reveals Three Sources of Adaptive Variation during a Rapid Radiation
/ref>


References

{{Taxonbar, from=Q3489171 cheesmaniae Endemic flora of Ecuador Endemic flora of the Galápagos Islands Tomatoes