Snow Camouflage
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Snow camouflage is the use of a coloration or pattern for effective
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
in winter, often combined with a different summer camouflage. Summer patterns are typically
disruptively patterned Disruptive coloration (also known as disruptive camouflage or disruptive patterning) is a form of camouflage that works by breaking up the outlines of an animal, soldier or military hardware with a strongly contrasting pattern. It is often com ...
combinations of shades of browns and greys, up to black, while winter patterns are dominated by white to match snowy landscapes. Among animals, variable snow camouflage is a type of
seasonal polyphenism A polyphenic trait is a trait for which multiple, discrete phenotypes can arise from a single genotype as a result of differing environmental conditions. It is therefore a special case of phenotypic plasticity. There are several types of polyphen ...
with a distinct winter
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
or
pelage A fur is a soft, thick growth of hair that covers the skin of almost all mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an ...
. It is found in birds such as the
rock ptarmigan The rock ptarmigan (''Lagopus muta'') is a medium-sized game bird in the grouse family. It is known simply as the ptarmigan in Europe. It is the official bird for the Canadian territory of Nunavut, where it is known as the ''aqiggiq'' (ᐊᕿ ...
,
lagomorphs The lagomorphs () are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae (rabbits and hares) and the Ochotonidae (pikas). There are 110 recent species of lagomorph, of which 109 species in twelve ...
such as the
Arctic hare The Arctic hare (''Lepus arcticus'') is a species of hare highly adapted to living in the Arctic tundra and other icy biomes. The Arctic hare survives with shortened ears and limbs, a small nose, fat that makes up close to 20% of its body, and a ...
,
mustelids The Mustelidae (; from Latin , weasel) are a diverse family of carnivoran mammals, including weasels, badgers, otters, polecats, martens, grisons, and wolverines. Otherwise known as mustelids (), they form the largest family in the suborder Can ...
such as the
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine or ermine, is a species of mustelid native to Eurasia and the northern regions of North America. Because of its wide circumpolar distribution, it is listed as Least Concern on th ...
, and one
canid Canidae (; from Latin, ''canis'', "dog") is a family (biology), biological family of caniform carnivorans, constituting a clade. A member of this family is also called a canid (). The family includes three subfamily, subfamilies: the Caninae, a ...
, the
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small species of fox native to the Arctic regions of the Northern Hemisphere and common throughout the Tundra#Arctic tundra, Arctic tundra biome. I ...
. Since these have evolved separately, the similar appearance is due to
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
. This was used as early evidence for natural selection. Some high Arctic species like the
snowy owl The snowy owl (''Bubo scandiacus''), also known as the polar owl, the white owl and the Arctic owl, is a large, white owl of the true owl family. Snowy owls are native to the Arctic regions of both North America and the Palearctic, breeding mo ...
and
polar bear The polar bear (''Ursus maritimus'') is a large bear native to the Arctic and nearby areas. It is closely related to the brown bear, and the two species can Hybrid (biology), interbreed. The polar bear is the largest extant species of bear ...
however remain white all year round. In military usage, soldiers often either exchange their disruptively-patterned summer uniforms for thicker snow camouflage uniforms printed with mainly-white versions of
camouflage patterns Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the ...
in winter, or they wear white overalls over their uniforms. Some armies have made use of reversible uniforms, printed in different seasonal patterns on their two sides. Vehicles and guns are often simply repainted in white. Occasionally, aircraft too are repainted in snow camouflage patterns.


Among animals


White as camouflage

Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
mentioned the white winter coloration of the ptarmigan in his 1859 ''
Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life'')The book's full original title was ''On the Origin of Species by M ...
'': The white protective coloration of arctic animals was noted by an early student of camouflage, the naturalist
Alfred Russel Wallace Alfred Russel Wallace (8 January 1823 – 7 November 1913) was an English naturalist, explorer, geographer, anthropologist, biologist and illustrator. He independently conceived the theory of evolution through natural selection; his 1858 pap ...
, in his 1889 book ''
Darwinism ''Darwinism'' is a term used to describe a theory of biological evolution developed by the English naturalist Charles Darwin (1809–1882) and others. The theory states that all species of organisms arise and develop through the natural sel ...
''; he listed the
polar bear The polar bear (''Ursus maritimus'') is a large bear native to the Arctic and nearby areas. It is closely related to the brown bear, and the two species can Hybrid (biology), interbreed. The polar bear is the largest extant species of bear ...
, the American polar hare, the
snowy owl The snowy owl (''Bubo scandiacus''), also known as the polar owl, the white owl and the Arctic owl, is a large, white owl of the true owl family. Snowy owls are native to the Arctic regions of both North America and the Palearctic, breeding mo ...
and the
gyr falcon The gyrfalcon ( or ) (), also abbreviated as gyr, is a bird of prey from the genus ''Falco'' (falcons and kestrels) and the largest species of the family Falconidae. A high-latitude species, the gyrfalcon breeds on the Arctic coasts and tundra, ...
as remaining white all year, while the arctic fox, arctic hare, ermine and ptarmigan change their colour, and observed "the obvious explanation", that it was for concealment, at a time when Darwinism was at a low ebb. Later zoologists such as Hugh B. Cott have echoed his observations, adding that other animals of the Arctic such as
musk ox Musk is a class of aromatic substances commonly used as base notes in perfumery. They include glandular secretions from animals such as the musk deer, numerous plants emitting similar fragrances, and artificial substances with similar odors. ' ...
,
moose The moose (: 'moose'; used in North America) or elk (: 'elk' or 'elks'; used in Eurasia) (''Alces alces'') is the world's tallest, largest and heaviest extant species of deer and the only species in the genus ''Alces''. It is also the tal ...
,
reindeer The reindeer or caribou (''Rangifer tarandus'') is a species of deer with circumpolar distribution, native to Arctic, subarctic, tundra, taiga, boreal, and mountainous regions of Northern Europe, Siberia, and North America. It is the only re ...
,
wolverine The wolverine ( , ; ''Gulo gulo''), also called the carcajou or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species, member of the family Mustelidae. It is a muscular carnivore and a solitary animal. The w ...
and
raven A raven is any of several large-bodied passerine bird species in the genus '' Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between crows and ravens; the two names are assigne ...
never become white "even in the coldest parts of their range". Cott noted that both animals that hunt, like polar bear and stoat, and prey animals like ptarmigan and mountain hare, require camouflage to hide from prey or from predators respectively. There is little experimental evidence for the adaptiveness of white as camouflage, though the ornithologist W. L. N. Tickell, reviewing proposed explanations of white plumage in birds, writes that in the ptarmigan "it is difficult to escape the conclusion that cryptic brown summer plumage becomes a liability in snow, and white plumage is therefore another cryptic adaptation." All the same, he notes, "in spite of winter plumage, many Ptarmigan in NE Iceland are killed by Gyrfalcons throughout the winter."


Seasonal polyphenism

Some animals of the far north, like the
snowshoe Snowshoes are specialized outdoor gear for walking over snow. Their large footprint spreads the user's weight out and allows them to travel largely on top of rather than through snow. Adjustable bindings attach them to appropriate winter footw ...
and
Arctic hare The Arctic hare (''Lepus arcticus'') is a species of hare highly adapted to living in the Arctic tundra and other icy biomes. The Arctic hare survives with shortened ears and limbs, a small nose, fat that makes up close to 20% of its body, and a ...
s,
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small species of fox native to the Arctic regions of the Northern Hemisphere and common throughout the Tundra#Arctic tundra, Arctic tundra biome. I ...
,
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine or ermine, is a species of mustelid native to Eurasia and the northern regions of North America. Because of its wide circumpolar distribution, it is listed as Least Concern on th ...
, and
rock ptarmigan The rock ptarmigan (''Lagopus muta'') is a medium-sized game bird in the grouse family. It is known simply as the ptarmigan in Europe. It is the official bird for the Canadian territory of Nunavut, where it is known as the ''aqiggiq'' (ᐊᕿ ...
change their coat colour (by moulting and growing new fur or feathers) from brown or grey summer
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
to white in the winter; the Arctic fox is the only species in the dog family to do so. However, Arctic hares which live in the far north of
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
, where summer is very short, remain white year-round. Since these animals in widely separated groups have evolved separately, the similarity of coloration is due to
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
, on the presumption that
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the Heredity, heritable traits characteristic of a population over generation ...
favours a particular coloration in a particular environment. The
seasonal polyphenism A polyphenic trait is a trait for which multiple, discrete phenotypes can arise from a single genotype as a result of differing environmental conditions. It is therefore a special case of phenotypic plasticity. There are several types of polyphen ...
in
willow grouse The willow ptarmigan ( ); ''Lagopus lagopus'') or willow grouse is a bird in the grouse subfamily Tetraoninae of the pheasant family Phasianidae. It is also known colloquially as awebo bird. The willow ptarmigan breeds in birch and other forest ...
differs between Scottish and Scandinavian populations. In Scotland, grouse have two plumages (breeding and non-breeding), while in Scandinavia there is a third plumage, a white winter morph. The genetic basis for this is not in the
melanin Melanin (; ) is a family of biomolecules organized as oligomers or polymers, which among other functions provide the pigments of many organisms. Melanin pigments are produced in a specialized group of cells known as melanocytes. There are ...
pigment system, and is probably due to regulatory changes. The behaviour of moulting females in springtime depends on their plumage state: they tend to sit on snow while they are mainly white, but choose the border between bare ground and snow when they have more dark feathers. They seem to be choosing the best compromise between camouflage and food quality. The
effects of climate change Effects of climate change are well documented and growing for Earth's natural environment and human societies. Changes to the climate system include an Instrumental temperature record, overall warming trend, Effects of climate change on the ...
can lead to a mismatch between the seasonal coat coloration of arctic animals such as
snowshoe hare The snowshoe hare (''Lepus americanus''), also called the varying hare or snowshoe rabbit, is a species of hare found in North America. It has the name "snowshoe" because of the large size of its hind feet. The animal's feet prevent it from sink ...
s with the increasingly snow-free landscape.


Military usage

The principle of varying coloration with the changing seasons has military applications.


First World War

In the
First World War World War I or the First World War (28 July 1914 – 11 November 1918), also known as the Great War, was a World war, global conflict between two coalitions: the Allies of World War I, Allies (or Entente) and the Central Powers. Fighting to ...
, firing and observation positions were hand-painted in disruptive patterns by artists known as
camoufleurs A camoufleur or camouflage officer is a person who designed and implemented military camouflage in one of the world wars of the twentieth century. The term originally meant a person serving in a First World War French military camouflage unit. In ...
, and they sometimes varied their patterns seasonally. Uniforms were largely of a single colour, such as the British
khaki The color khaki (, ) is a light shade of tan (color), tan with a slight yellowish tinge. Khaki has been used by many armies around the world for uniforms and equipment, particularly in arid or desert regions, where it provides camouflage rela ...
; but snow camouflage clothes came into use in some armies as the war progressed. For example, Austro-Hungarian troops on the Italian front used
ski Skis are runners, attached to the user's feet, designed to glide over snow. Typically employed in pairs, skis are attached to ski boots with ski bindings, with either a free, lockable, or partially secured heel. For climbing slopes, ski skins c ...
s and wore snow camouflage smocks and overtrousers over their
uniforms A uniform is a variety of costume worn by members of an organization while usually participating in that organization's activity. Modern uniforms are most often worn by armed forces and paramilitary organizations such as police, emergency ser ...
, and improvised white cloths over their uniform caps.


Second World War

Several armies in the
Second World War World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...
in Northern European countries preferred separate winter uniforms rather than oversuits. The
Waffen-SS The (; ) was the military branch, combat branch of the Nazi Party's paramilitary ''Schutzstaffel'' (SS) organisation. Its formations included men from Nazi Germany, along with Waffen-SS foreign volunteers and conscripts, volunteers and conscr ...
went a step further, developing reversible uniforms with separate schemes for summer and autumn, as well as white winter oversuits. Other German units fighting in Eastern Europe were at first poorly equipped for winter, having to make do with ordinary summer uniforms, but in the winter of 1942 to 1943 new white two-piece hooded oversuits with long mitten gauntlets started to arrive. American troops in Europe in the winter of 1944 to 1945 improvised snow capes and helmet covers from white cloth such as bed linen. The Red Army issued a report, "Tactical and Technical trends, No. 17" in January 1943 on the camouflage of tanks in winter. It advised either all-white using zinc white or titanium white paint for level, open country, or disruptive two-colour winter camouflage for areas with more variety including "forests, underbrush, small settlements, thawed patches of earth". The two colours could be achieved either by leaving around a quarter to a third of the vehicle's summer camouflage uncovered, or by repainting the whole vehicle in white with dark gray or gray-brown spots. Units were advised not to paint all their vehicles identically, but to have some tanks all white, some in white with green stripes, and some in white and gray or gray-brown. Winter camouflage was not limited to paint: tracks left in the snow were to be obliterated, vehicles parked in cover, headlights covered with white fabric, shelters constructed, or else vehicles covered with white fabric, or dark fabric scattered with snow.


Post-war

Later in the 20th century, as
disruptively patterned Disruptive coloration (also known as disruptive camouflage or disruptive patterning) is a form of camouflage that works by breaking up the outlines of an animal, soldier or military hardware with a strongly contrasting pattern. It is often com ...
uniforms became widespread, winter variants of patterns such as ''
Flecktarn ''Flecktarn'' (; "mottled camouflage"; also known as ''Flecktarnmuster'' or ''Fleckentarn'') is a family of three-, four-, five- or six-color disruptive camouflage patterns, the most common being the five-color pattern, consisting of dark gree ...
'' were developed, the background colour (such as green) being replaced with white to form a ''Schneetarn'' (snow camouflage pattern). Aircraft deployed in winter have sometimes been snow camouflaged, as with
Sepecat Jaguar The SEPECAT Jaguar is a British-French supersonic jet attack aircraft originally used by the British Royal Air Force and the French Air Force in the close air support and nuclear strike role. As of 2025, the Jaguar remains in service with the ...
s on exercise in Norway.


References

{{Camouflage Camouflage mechanisms Animal coat colors Military camouflage types