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A slough ( or ) is a
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
, usually a
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
or shallow
lake A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from ...
, often a backwater to a larger body of water. Water tends to be stagnant or may flow slowly on a seasonal basis. In North America, "slough" may refer to a side-channel from or feeding a river, or an inlet or natural channel only sporadically filled with water. An example of this is Finn Slough on the
Fraser River The Fraser River () is the longest river within British Columbia, Canada, rising at Fraser Pass near Blackrock Mountain (Canada), Blackrock Mountain in the Rocky Mountains and flowing for , into the Strait of Georgia just south of the City of V ...
, whose lower reaches have dozens of notable sloughs. Some sloughs, like Elkhorn Slough, used to be mouths of rivers, but have become stagnant because tectonic activity cut off the river's source. In the
Sacramento River The Sacramento River () is the principal river of Northern California in the United States and is the largest river in California. Rising in the Klamath Mountains, the river flows south for before reaching the Sacramento–San Joaquin River D ...
, Steamboat Slough was an alternate branch of the river, a preferred shortcut route for
steamboat A steamboat is a boat that is marine propulsion, propelled primarily by marine steam engine, steam power, typically driving propellers or Paddle steamer, paddlewheels. The term ''steamboat'' is used to refer to small steam-powered vessels worki ...
s passing between
Sacramento Sacramento ( or ; ; ) is the capital city of the U.S. state of California and the seat of Sacramento County. Located at the confluence of the Sacramento and American Rivers in Northern California's Sacramento Valley, Sacramento's 2020 p ...
and
San Francisco San Francisco, officially the City and County of San Francisco, is a commercial, Financial District, San Francisco, financial, and Culture of San Francisco, cultural center of Northern California. With a population of 827,526 residents as of ...
. Georgiana Slough was a steamboat route through the
Sacramento–San Joaquin River Delta The Sacramento–San Joaquin River Delta, or California Delta, is an expansive inland river delta and estuary in Central California and Northern California Northern California (commonly shortened to NorCal) is a geocultural region that ...
, from the Sacramento River to the San Joaquin River and Stockton.


Plants and animals

A slough, also called a tidal channel, is a channel in a
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
. Typically, it is either stagnant or slow flowing on a seasonal basis. Vegetation patterns in a slough are largely determined by depth, distribution, and duration in the environment. Moreover, these same variables also influence the distribution, abundance, reproduction, and seasonal movements of aquatic and terrestrial life within the sloughs. Sloughs support a wide variety of plant life that is adapted to rapidly changing physical conditions such as
salinity Salinity () is the saltiness or amount of salt (chemistry), salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensio ...
, oxygen levels and depth. In general, sloughs are microhabitats high in species diversity. Open water sloughs are characterized by submerged and floating vegetation which includes
periphyton Periphyton is a complex mixture of algae, cyanobacteria, heterotrophic microbes, and detritus that is attached to submerged surfaces in most aquatic ecosystems. The related term Aufwuchs ( German "surface growth" or "overgrowth", ) refers to the ...
mats dominated by sawgrass typically. The topographical and vegetation heterogeneity of ridge and slough landscape influences the productivity and diversity of birds and fish adapted to that wetland. Fish that typically inhabit sloughs include tidewater goby, California killifish, mosquitofish, and topsmelt. Food habits of fish within sloughs consist of preying upon
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s; mostly epifaunal
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
followed by epifaunal and infaunal worms and mollusks. Fish can feed on zooplankton and plant material. Research on prey species for fish in sloughs found that in a study done on Elkhorn Slough in California the mean prey richness for fish was greatest near the ocean and lowest inshore. This allows for a higher availability of food to enhance the function of inshore habitats and emphasizes the importance of invertebrate prey populations and how they influence plant production.
Bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s also inhabit sloughs, making them hotspots for
birdwatching Birdwatching, or birding, is the observing of birds, either as a recreational activity or as a form of citizen science. A birdwatcher may observe by using their naked eye, by using a visual enhancement device such as binoculars or a telescop ...
, with the Elkhorn Slough being one of the premier birdwatching sites in the western United States. Over 340 species have been seen visiting, including several rare and endangered species. Bird species seen in sloughs include acorn woodpecker, brown pelican, Caspian tern,
great blue heron The great blue heron (''Ardea herodias'') is a large wading bird in the heron family Ardeidae, common near the shores of open water and in wetlands over most of North and Central America, as well as far northwestern South America, the Caribbea ...
,
great egret The great egret (''Ardea alba''), also known as the common egret, large egret, great white egret, or great white heron, is a large, widely distributed egret. The four subspecies are found in Asia, Africa, the Americas, and southern Europe. R ...
,
great horned owl The great horned owl (''Bubo virginianus''), also known as the tiger owl (originally derived from early naturalists' description as the "winged tiger" or "tiger of the air") or the hoot owl, is a large owl native to the Americas. It is an extreme ...
,
snowy plover The snowy plover (''Anarhynchus nivosus'') is a small shorebird found in the Americas. It is a member of the bird family Charadriidae, which includes the plovers, dotterels, and lapwings. The snowy plover was originally described by John Cassin ...
, and white-tailed kite. Sloughs are largely influenced by human development such as urban and
agricultural expansion Agricultural expansion describes the growth of agricultural land ( arable land, pastures, etc.) especially in the 20th and 21st centuries. The agricultural expansion is often explained as a direct consequence of the global increase in food and e ...
, industrial and agricultural practices, water management practices, and humans influence on
species composition Relative species abundance is a component of biodiversity and is a measure of how common or rare a species is relative to other species in a defined location or community.Hubbell, S. P. 2001. ''The unified neutral theory of biodiversity and biogeog ...
. Uses of identifying these aspects of human involvement can help to better predict restoration efforts to be made in managing sloughs. Examples of attributes that are affected by human stress upon the environment include
periphyton Periphyton is a complex mixture of algae, cyanobacteria, heterotrophic microbes, and detritus that is attached to submerged surfaces in most aquatic ecosystems. The related term Aufwuchs ( German "surface growth" or "overgrowth", ) refers to the ...
, marsh plant communities, tree islands,
alligator An alligator, or colloquially gator, is a large reptile in the genus ''Alligator'' of the Family (biology), family Alligatoridae in the Order (biology), order Crocodilia. The two Extant taxon, extant species are the American alligator (''A. mis ...
s, wading birds, and marsh fishes,
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s, and herpetofauna.


Formation

A slough can form when a
meander A meander is one of a series of regular sinuous curves in the Channel (geography), channel of a river or other watercourse. It is produced as a watercourse erosion, erodes the sediments of an outer, concave bank (cut bank, cut bank or river cl ...
gets cut off from the main river channel creating an
oxbow lake An oxbow lake is a U-shaped lake or stream pool, pool that forms when a wide meander of a river is meander cutoff, cut off, creating a free-standing body of water. The word "oxbow" can also refer to a U-shaped bend in a river or stream, whether ...
that accumulates with fine overbank sediment and organic material such as
peat Peat is an accumulation of partially Decomposition, decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, Moorland, moors, or muskegs. ''Sphagnum'' moss, also called peat moss, is one of the most ...
. This creates a
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
or
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
environment. One end of the oxbow configuration then continues to receive flow from the main channel, creating a slough. Sloughs are typically associated with the ridge formations found in their presence. Such a landscape consists of mosaic linear ridges, typically of some sort of grass such as sawgrass ridges in the Florida
Everglades The Everglades is a natural region of flooded grasslands in the southern portion of the U.S. state of Florida, comprising the southern half of a large drainage basin within the Neotropical realm. The system begins near Orlando with the K ...
, that are separated by deeper water sloughs. Edges of sloughs are layers of
sediment Sediment is a solid material that is transported to a new location where it is deposited. It occurs naturally and, through the processes of weathering and erosion, is broken down and subsequently sediment transport, transported by the action of ...
deposited by a river over time. The development of this landscape is thought to occur by the preferential formation of peat in
bedrock In geology, bedrock is solid rock that lies under loose material ( regolith) within the crust of Earth or another terrestrial planet. Definition Bedrock is the solid rock that underlies looser surface material. An exposed portion of bed ...
depressions. Multiple of these deposits mounted on top of the surrounding bedrock can become elongated alongside the slough and create flow diversions within the system. Different rates of this
peat Peat is an accumulation of partially Decomposition, decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, Moorland, moors, or muskegs. ''Sphagnum'' moss, also called peat moss, is one of the most ...
accumulation could be triggered by variations in microtopography that alter plant production and vegetation type. Water flow might be the key to preventing an accumulation of organic sediment in sloughs due to the fact that accumulation leads to lowering water depths and instead allows for the growth of vegetation.


Degradation

Overall little quantitative data on the degradation of slough landscape exists. Slough and ridge landscape has been greatly degraded in terms of both topographic and vegetation changes over time. Topographical changes create an increase in the relief between ridge crests and slough bottoms. Vegetation changes consist of an increase in the amount of dense grass and decrease in the area of open water, creating a blurring of the directional ridge and slough pattern. Historical everglade and slough landscape has been greatly affected and degraded by human activity. Open water sloughs support important ecological functions that have been seen to be sensitive to hydrologic and water quality problems stemmed from human activities.


Restoration

Sloughs are ecologically important as they are a part of an
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
environment: wetlands. They act as a buffer from land to sea and act as an active part of the estuary system where freshwater flows from creeks and runoff from the land mix with salty ocean water transported by tides. Restoration is a big effort in California wetlands to restore slough and ridge landscapes. Examples of restoration projects on slough landscapes include The Elkhorn Slough Tidal Wetland Project, Dutch Slough Tidal Restoration Project, and the McDaniel Slough wetland enhancement project.


See also


References

{{Wetlands Hydrology Wetlands River morphology