
Shrubland, scrubland, scrub, brush, or bush is a
plant community
A plant community is a collection or association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The components of each plant ...
characterized by
vegetation
Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic charact ...
dominated by
shrubs, often also including
grass
Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in law ...
es,
herb
In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicina ...
s, and
geophyte
A storage organ is a part of a plant specifically modified for storage of energy
(generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have ...
s. Shrubland may either occur naturally or be the result of human activity. It may be the mature
vegetation type
Vegetation classification is the process of classifying and mapping the vegetation over an area of the earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource and environmental management ...
in a particular region and remain stable over time, or a transitional community that occurs temporarily as the result of a disturbance, such as fire. A stable state may be maintained by regular natural disturbance such as fire or
browsing
Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
. Shrubland may be unsuitable for human habitation because of the danger of fire. The term was coined in 1903.
Shrubland species generally show a wide range of
adaptations
In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
to fire, such as heavy seed production,
lignotubers, and fire-induced germination.
Botanical structural form
In
botany
Botany, also called plant science (or plant sciences), plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "bot ...
and
ecology
Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overl ...
a shrub is defined as a much-branched
woody plant
A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposite to herbaceous plants that die back to the ground until s ...
less than 8 m high and usually with many
stems. Tall shrubs are mostly 2–8 m high, small shrubs 1–2 m high and
subshrub
A subshrub ( Latin ''suffrutex'') or dwarf shrub is a short shrub, and is a woody plant. Prostrate shrub is a related term. "Subshrub" is often used interchangeably with "bush".Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their ...
s less than 1 m high.
There is a descriptive system widely adopted in
Australia to describe different types of vegetation is based on
structural characteristics based on
plant life-form
Plant life-form schemes constitute a way of classifying plants alternatively to the ordinary species-genus-family scientific classification. In colloquial speech, plants may be classified as trees, shrubs, herbs (forbs and graminoids), etc. The sc ...
, plus the height and foliage cover of the tallest stratum or
dominant species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
.

For shrubs 2–8 m high the following structural forms result:
* dense foliage cover (70–100%) — closed-scrub
* mid-dense foliage cover (30–70%) — open-scrub
* very sparse foliage cover (<10%) — tall open shrubland
For shrubs <2 m high the following structural forms result:
* dense foliage cover (70–100%) — closed-
heath
A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a cooler ...
* mid-dense foliage cover (30–70%) — open-heath
* sparse foliage cover (10–30%) — low shrubland
* very sparse foliage cover (<10%) — low open shrubland
Biome plant group

Similarly, shrubland is a category used to describe a type of
biome
A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
plant group. In this context, shrublands are dense thickets of evergreen
sclerophyll
Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...
shrubs and small trees,
called:
*
Chaparral
Chaparral ( ) is a shrubland plant community and geographical feature found primarily in the U.S. state of California, in southern Oregon, and in the northern portion of the Baja California Peninsula in Mexico. It is shaped by a Mediterrane ...
in
California
California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
*
Matorral
300px, Springtime in Chilean matorral a few kilometers north of Santiago along the Pan-American Highway
Matorral is a Spanish language, Spanish word, along with ''tomillares'', for shrubland, thicket or bushes. It is used in naming and describin ...
in
Chile
Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
,
Mexico
Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
, and
Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond")
, national_anthem = (English: "Royal March")
, ...
*
Maquis
Maquis may refer to:
Resistance groups
* Maquis (World War II), predominantly rural guerrilla bands of the French Resistance
* Spanish Maquis, guerrillas who fought against Francoist Spain in the aftermath of the Spanish Civil War
* The network ...
in
France
France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan ar ...
and elsewhere around the
Mediterranean
The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on th ...
*
Macchia in
Italy
Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
*
Fynbos in
South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
*
in
Sydney,
Australia
*
Kwongan
Kwongan is plant community found in south-western Western Australia. The name is a Bibbelmun (Noongar) Aboriginal term of wide geographical use defined by Beard (1976) as
Kwongan has replaced other terms applied by European botanists such as ...
in Southwest
Australia
*
Cedar scrub in
Texas Hill Country
The Texas Hill Country is a geographic region of Central and South Texas, forming the southeast part of the Edwards Plateau. Given its location, climate, terrain, and vegetation, the Hill Country can be considered the border between the Amer ...
In some places shrubland is the mature vegetation type, and in other places the result of degradation of former
forest
A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
or
woodland
A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
by
logging or
overgrazing
Overgrazing occurs when plants are exposed to intensive grazing for extended periods of time, or without sufficient recovery periods. It can be caused by either livestock in poorly managed agricultural applications, game reserves, or nature ...
, or disturbance by major fires.
A number of
World Wildlife Fund
The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wor ...
biomes are characterized as shrublands, including:
[
;Desert scrublands
]
''Xeric'' or ''desert scrublands'' occur in the world's deserts and xeric shrublands
Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (ancient Greek xērós, “dry") shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in thi ...
ecoregions, or in areas of fast-draining sandy soils in more humid regions. These scrublands are characterized by plants with adaptations to the dry climate, which include small leaves to limit water loss, thorns to protect them from grazing animals, succulent leaves or stems, storage organ
A storage organ is a part of a plant specifically modified for storage of energy
(generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have ...
s to store water, and long taproots to reach groundwater.
;Mediterranean scrublands
''Mediterranean scrublands'' occur naturally in the Mediterranean forests, woodlands, and scrub
Mediterranean forests, woodlands, and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in ...
biome
A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
s, located in the five Mediterranean climate
A Mediterranean climate (also called a dry summer temperate climate ''Cs'') is a temperate climate sub-type, generally characterized by warm, dry summers and mild, fairly wet winters; these weather conditions are typically experienced in the ...
regions of the world. Scrublands are most common near the seacoast, and have often adapted to the wind and salt air of the ocean. Low, soft-leaved scrublands around the Mediterranean Basin
In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and w ...
are known as ''garrigue
Garrigue or garigue ( ), also known as phrygana ( el, φρύγανα , n. pl.), is a type of low scrubland ecoregion and plant community in the Mediterranean forests, woodlands, and scrub biome.
It is found on limestone soils in southern Fran ...
'' in France
France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan ar ...
, ''phrygana'' in Greece
Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wit ...
, ''tomillares'' in Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond")
, national_anthem = (English: "Royal March")
, ...
, and ''batha'' in Israel
Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
. Northern coastal scrub and coastal sage scrub
Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is ...
occur along the California
California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
coast, strandveld in the Western Cape
The Western Cape is a province of South Africa, situated on the south-western coast of the country. It is the fourth largest of the nine provinces with an area of , and the third most populous, with an estimated 7 million inhabitants in 2020 ...
of South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
, coastal matorral in central Chile
Central Chile (''Zona central'') is one of the five natural regions into which CORFO divided continental Chile in 1950. It is home to a majority of the Chilean population and includes the three largest metropolitan areas—Santiago, Valparaí ...
, and sand-heath and kwongan
Kwongan is plant community found in south-western Western Australia. The name is a Bibbelmun (Noongar) Aboriginal term of wide geographical use defined by Beard (1976) as
Kwongan has replaced other terms applied by European botanists such as ...
in Southwest Australia
Southwest Australia is a biogeographic region in Western Australia. It includes the Mediterranean-climate area of southwestern Australia, which is home to a diverse and distinctive flora and fauna.
The region is also known as the Southwest Aus ...
.
;Interior scrublands
''Interior scrublands'' occur naturally in semi-arid areas where soils are nutrient-poor, such as on the matas of Portugal
Portugal, officially the Portuguese Republic, In recognized minority languages of Portugal:
:* mwl, República Pertuesa is a country located on the Iberian Peninsula, in Southwestern Europe, and whose territory also includes the Macaronesian ...
which are underlain by Cambrian
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
and Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoi ...
schists. Florida scrub
Florida sand pine scrub is an endangered subtropical forest ecoregion found throughout Florida in the United States. It is found on coastal and inland sand ridges and is characterized by an evergreen xeromorphic plant community dominated by shrub ...
is another example of interior scrublands.
;Dwarf shrubs
Some vegetation types are formed of '' dwarf-shrubs'': low-growing or creeping shrubs. These include the maquis
Maquis may refer to:
Resistance groups
* Maquis (World War II), predominantly rural guerrilla bands of the French Resistance
* Spanish Maquis, guerrillas who fought against Francoist Spain in the aftermath of the Spanish Civil War
* The network ...
and garrigue
Garrigue or garigue ( ), also known as phrygana ( el, φρύγανα , n. pl.), is a type of low scrubland ecoregion and plant community in the Mediterranean forests, woodlands, and scrub biome.
It is found on limestone soils in southern Fran ...
s of Mediterranean climates, and the acid-loving dwarf shrubs of heathland
A heath () is a shrubland habitat (ecology), habitat found mainly on free-draining infertile, acidic soils and characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great B ...
and moorland.
See also
* Fynbos
* Maquis
Maquis may refer to:
Resistance groups
* Maquis (World War II), predominantly rural guerrilla bands of the French Resistance
* Spanish Maquis, guerrillas who fought against Francoist Spain in the aftermath of the Spanish Civil War
* The network ...
* Prostrate shrub
A prostrate shrub is a woody plant, most of the branches of which lie upon or just above the ground, rather than being held erect as are the branches of most trees and shrubs.
Background
Prostration may occur because the supporting tissues in ste ...
* Semi-desert
A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-a ...
* Shrub-steppe
* Shrub swamp
Shrub swamps — also called scrub swamps or buttonbush swamps — are a type of freshwater wetland ecosystem
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These b ...
* Moorland
Notes and references
External links
*
{{Vegetation