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__NOTOC__ Mrs. Ples is the popular nickname for the most complete skull of an ''
Australopithecus africanus ''Australopithecus africanus'' is an extinct species of australopithecine which lived between about 3.3 and 2.1 million years ago in the Late Pliocene to Early Pleistocene of South Africa. The species has been recovered from Taung, Sterkfontei ...
'' ever found in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. Many ''Australopithecus'' fossils have been found near
Sterkfontein Sterkfontein (Afrikaans for ''Strong Spring'') is a set of limestone caves of special interest in paleoanthropology located in Gauteng province, about northwest of Johannesburg, South Africa in the Muldersdrift area close to the town of K ...
, about northwest of
Johannesburg Johannesburg ( , , ; Zulu language, Zulu and Xhosa language, Xhosa: eGoli ) (colloquially known as Jozi, Joburg, Jo'burg or "The City of Gold") is the most populous city in South Africa. With 5,538,596 people in the City of Johannesburg alon ...
, in a region of
Gauteng Gauteng ( , ; Sotho-Tswana languages, Sotho-Tswana for 'place of gold'; or ) is one of the nine provinces of South Africa. Situated on the Highveld, Gauteng is the smallest province by land area in South Africa. Although Gauteng accounts f ...
(part of the old Transvaal) now designated as the
Cradle of Humankind The Cradle of Humankind is a paleoanthropological site that is located about northwest of Johannesburg, South Africa, in the Gauteng province. Declared a World Heritage Site by UNESCO in 1999, the site is home to the largest known concentrat ...
World Heritage Site World Heritage Sites are landmarks and areas with legal protection under an treaty, international treaty administered by UNESCO for having cultural, historical, or scientific significance. The sites are judged to contain "cultural and natural ...
. Mrs. Ples was discovered by
Robert Broom Robert Broom Fellow of the Royal Society, FRS FRSE (30 November 1866 6 April 1951) was a British- South African medical doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University ...
and
John T. Robinson John Talbot Robinson FRSSAf (10 January 1923 – 12 October 2001) was a distinguished South African hominin paleontologist. His most famous discovery (with Robert Broom) was the nearly complete fossil skull of the hominin species ''Australop ...
on April 18, 1947. Because of Broom's use of dynamite and pickaxe while excavating, Mrs. Ples's skull was blown into pieces and some fragments are missing. Nonetheless, Mrs./Mr. Ples is one of the most "perfect" pre-human skulls ever found. The skull is currently held at the
Ditsong National Museum of Natural History The Ditsong National Museum of Natural History, formerly the Transvaal Museum, is a natural history museum situated in Pretoria, South Africa. It is located on Paul Kruger Street, between Visagie and Minnaar Streets, opposite the Pretoria City H ...
in
Pretoria Pretoria ( ; ) is the Capital of South Africa, administrative capital of South Africa, serving as the seat of the Executive (government), executive branch of government, and as the host to all foreign embassies to the country. Pretoria strad ...
. The nickname "Mrs. Ples" was coined by Broom's young co-workers. It derives from the scientific name ''Plesianthropus transvaalensis'' (near-man from the Transvaal), that Broom initially gave the skull, later subsumed ( synonymized) into the species ''Australopithecus africanus.'' In scientific publications, the specimen is referred to by its catalogue number, STS 5. The genus ''
Australopithecus ''Australopithecus'' (, ; or (, ) is a genus of early hominins that existed in Africa during the Pliocene and Early Pleistocene. The genera ''Homo'' (which includes modern humans), ''Paranthropus'', and ''Kenyanthropus'' evolved from some ''Aus ...
'', of which there are several species, is considered the likely precursor of the genus ''
Homo ''Homo'' () is a genus of great ape (family Hominidae) that emerged from the genus ''Australopithecus'' and encompasses only a single extant species, ''Homo sapiens'' (modern humans), along with a number of extinct species (collectively called ...
''. Though its
cranium The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
is comparable to a
chimpanzee The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
's, ''Australopithecus'' walked upright, as humans do. This was a surprise to anthropologists at the time, because it had been assumed that the big brain of ''Homo'' had preceded, or at least evolved in tandem with, our upright gait. Mrs. Ples, whose cranial capacity is only about , was one of the first fossils to reveal that upright walking had evolved well before any significant growth in brain size. The sex of the specimen is not completely certain and so ''Mrs. Ples'' may in fact be ''Mr. Ples''. Moreover, X-ray analysis of the specimen's teeth (''see below'') has suggested it was an adolescent. Hence a designation of ''
Miss Miss (pronounced ) is an English-language honorific typically used for a girl, for an unmarried woman (when not using another title such as " Doctor" or "Dame"), or for a married woman retaining her maiden name. Originating in the 17th century ...
Ples'' or ''
Master Master, master's or masters may refer to: Ranks or titles In education: *Master (college), head of a college *Master's degree, a postgraduate or sometimes undergraduate degree in the specified discipline *Schoolmaster or master, presiding office ...
Ples'' is also possible. The paleoanthropologist Prof. Frederick E. Grine has studied the dental morphology of Mrs./Mr. Ples with a view to finally establishing Mrs./Ms. Ples's sex. Using the Computed Tomography (CT) scans of STS 5 from the experiments of Weber et al., (2012) they compared them to CT scans of more recently discovered ''A. africanus'' skulls from Sterkfontein. These scans allowed Grine to reconstruct the roots of the teeth, in order to see how the molar and canine teeth developed. This study concluded that Mrs./Mr. Ples was indeed a middle-aged female. However, the question is not entirely settled, since other studies have come to the opposite conclusion. Some experts have suggested that a partial skeleton, known only by its catalogue number of STS 14, which was discovered in the same year, in the same geological deposit and in proximity to Mrs. Ples, may belong to this skull. If correct, this would make Mrs. Ples the South African counterpart to the famous
Lucy Lucy is an English language, English feminine given name derived from the Latin masculine given name Lucius with the meaning ''as of light'' (''born at dawn or daylight'', maybe also ''shiny'', or ''of light complexion''). Alternative spellings ar ...
fossil. This skull, along with others discovered at
Taung Taung is a small town situated in the North West Province of South Africa. The name means ''place of the lion'' and was named after Tau, the King of the Barolong people. ''Tau'' is the Tswana word for lion. Taung skull fossil site In 1924, a sk ...
, Sterkfontein, and
Makapansgat Makapansgat () (or Makapan Valley World Heritage Site) is an archaeological location within the Makapansgat and Zwartkrans Valleys, northeast of Mokopane in Limpopo province, South Africa. It is an important palaeontological site, with the loca ...
offered compelling evidence in favour of Charles Darwin's hypothesis that humanity's origin lay in Africa. The fossil was originally dated by a combination of
palaeomagnetism Paleomagnetism (occasionally palaeomagnetism) is the study of prehistoric Earth's magnetic fields recorded in rocks, sediment, or archeological materials. Geophysicists who specialize in paleomagnetism are called ''paleomagnetists.'' Certain ...
and uranium-lead techniques to 2.1-2.6 million years ago. A 2022 study dating surrounding rocks with a new methodology, stated their age as 3.4-3.7 million years old. But a more recent study, using the size of the teeth of ''Theropithecus oswaldi'', concluded that there is no evidence that any of the South African cave sites are older than about 2.8 million years. In 2004, Mrs. Ples was voted 95th in the
SABC 3 SABC 3, also branded as S3, is a South African free-to-air television channel owned by the South African Broadcasting Corporation (SABC). Since March 2024, it carries programming in English and Afrikaans only. History On 1 January 1982, two tel ...
's and
e.tv e.tv (commonly referred to on-air as e) is the first and only privately owned free-to-air television station in South Africa. Founded in 1998, the company is headquartered in Cape Town. It is the fifth terrestrial television channel in the ...
's '' Great South Africans'' Top 100 list.


See also

*
List of human evolution fossils The following tables give an overview of notable finds of Hominini, hominin fossils and Skeleton, remains relating to human evolution, beginning with the formation of the tribe Hominini (the divergence of the Chimpanzee–human last common ancest ...
*
Taung Child The Taung Child (or Taung Baby) is the fossilised skull of a young ''Australopithecus africanus''. It was discovered in 1924 by quarrymen working for the Northern Lime Company in Taung, South Africa. Raymond Dart described it as a new species ...


References


Further reading

*


External links


Maropeng - The Cradle of Humankind
official website
UNESCO - Fossil Hominid Sites of Sterkfontein, Swartkrans, Kromdraai, and EnvironsMetadata of STS 5 on NESPOS
{{Cradle of Humankind Australopithecus fossils Quaternary fossil record 1947 in paleontology Fossils of South Africa Archaeology of South Africa Cradle of Humankind