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A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a
rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...
or underground stem. In
grafting Grafting or graftage is a horticulture, horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the roots ...
, it refers to a plant, sometimes just a stump, which already has an established, healthy
root system In mathematics, a root system is a configuration of vector space, vectors in a Euclidean space satisfying certain geometrical properties. The concept is fundamental in the theory of Lie groups and Lie algebras, especially the classification and ...
, onto which a cutting or a bud from another plant is grafted. In some cases, such as vines of grapes and other berries, cuttings may be used for rootstocks, the roots being established in nursery conditions before planting them out. The plant part grafted onto the rootstock is usually called the scion. The scion is the plant that has the properties that
propagator In quantum mechanics and quantum field theory, the propagator is a function that specifies the probability amplitude for a particle to travel from one place to another in a given period of time, or to travel with a certain energy and momentum. I ...
desires above ground, including the photosynthetic activity and the fruit or decorative properties. The rootstock is selected for its interaction with the
soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...
, providing the roots and the stem to support the new plant, obtaining the necessary soil water and minerals, and resisting the relevant pests and diseases. After a few weeks, the tissues of the two parts will have grown together, eventually forming a single plant. After some years, it may be difficult to detect the site of the graft although the product always contains the components of two genetically different plants. The use of rootstocks is most commonly associated with
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
ing plants and trees, and is useful for mass propagating many other types of plants that do not breed true from
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
, or are particularly susceptible to
disease A disease is a particular abnormal condition that adversely affects the structure or function (biology), function of all or part of an organism and is not immediately due to any external injury. Diseases are often known to be medical condi ...
when grown on their own roots. Although grafting has been practiced for many hundreds of years, even in Roman times, most
orchard An orchard is an intentional plantation of trees or shrubs that is maintained for food production. Orchards comprise fruit tree, fruit- or nut (fruit), nut-producing trees that are generally grown for commercial production. Orchards are also so ...
rootstocks in current use were developed in the 20th century. A variety of rootstocks may be used for a single
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
or
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
of scion because different rootstocks impart different properties, such as vigour, fruit size and precocity. Rootstocks also may be selected for traits such as resistance to
drought A drought is a period of drier-than-normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, ...
, root pests, and
disease A disease is a particular abnormal condition that adversely affects the structure or function (biology), function of all or part of an organism and is not immediately due to any external injury. Diseases are often known to be medical condi ...
s.
Grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began approximately 8,0 ...
vines for commercial planting are most often grafted onto rootstocks to avoid damage by
phylloxera Grape phylloxera is an insect pest of grapevines worldwide, originally native to eastern North America. Grape phylloxera (''Daktulosphaira vitifoliae'' (Fitch 1855) belongs to the family Phylloxeridae, within the order Hemiptera, bugs); orig ...
, though vines available for sale to back garden viticulturists may not be. The rootstock may be a different species from the scion, but as a rule it should be closely related, for example, many commercial
pear Pears are fruits produced and consumed around the world, growing on a tree and harvested in late summer into mid-autumn. The pear tree and shrub are a species of genus ''Pyrus'' , in the Family (biology), family Rosaceae, bearing the Pome, po ...
s are grown on
quince The quince (; ''Cydonia oblonga'') is the sole member of the genus ''Cydonia'' in the Malinae subtribe (which contains apples, pears, and other fruits) of the Rosaceae family. It is a deciduous tree that bears hard, aromatic bright golden-yel ...
rootstock. Grafting can also be done in stages; a closely related scion is grafted to the rootstock, and a less closely related scion is grafted to the first scion. Serial grafting of several scions may also be used to produce a tree that bears several different fruit
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
s, with the same rootstock taking up and distributing water and minerals to the whole system. Those with more than three varieties are known as 'family trees'. When it is difficult to match a plant to the soil in a certain field or orchard, growers may graft a scion onto a rootstock that is compatible with the soil. It may then be convenient to plant a range of ungrafted rootstocks to see which suit the growing conditions best; the fruiting characteristics of the scion may be considered later, once the most successful rootstock has been identified. Rootstocks are studied extensively and often are sold with a complete guide to their ideal soil and climate. Growers determine the pH, mineral content,
nematode The nematodes ( or ; ; ), roundworms or eelworms constitute the phylum Nematoda. Species in the phylum inhabit a broad range of environments. Most species are free-living, feeding on microorganisms, but many are parasitic. Parasitic worms (h ...
population,
salinity Salinity () is the saltiness or amount of salt (chemistry), salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensio ...
, water availability,
pathogen In biology, a pathogen (, "suffering", "passion" and , "producer of"), in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a Germ theory of d ...
load and
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is usually defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural ...
iness of their particular soil, and select a rootstock which is matched to it.
Genetic testing Genetic testing, also known as DNA testing, is used to identify changes in DNA sequence or chromosome structure. Genetic testing can also include measuring the results of genetic changes, such as RNA analysis as an output of gene expression, or ...
is increasingly common, and new cultivars of rootstock are always being developed.


Uses in agriculture


AxR1

AxR1 is a
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began approximately 8,0 ...
rootstock once widely used in
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
viticulture Viticulture (, "vine-growing"), viniculture (, "wine-growing"), or winegrowing is the cultivation and harvesting of grapes. It is a branch of the science of horticulture. While the native territory of ''Vitis vinifera'', the common grape vine ...
. Its name is an abbreviation for "Aramon Rupestris Ganzin No. 1", which in turn is based on its parentage: a cross (made by a French grape hybridizer named Ganzin) between Aramon, a ''
Vitis vinifera ''Vitis vinifera'', the common grape vine, is a species of flowering plant, native to the Mediterranean Basin, Mediterranean region, Central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern ...
'' cultivar, and Rupestris, an American grape species, '' Vitis rupestris''also used on its own as rootstock, "Rupestris St. George" or "St. George," referring to a town in the South of France, Saint Georges d'Orques, where it was popular. It achieved a degree of notoriety in California when, after decades of recommendation as a preferred rootstockdespite repeated warnings from France and South Africa about its susceptibility (it had failed in Europe in the early 1900s)it ultimately succumbed to
phylloxera Grape phylloxera is an insect pest of grapevines worldwide, originally native to eastern North America. Grape phylloxera (''Daktulosphaira vitifoliae'' (Fitch 1855) belongs to the family Phylloxeridae, within the order Hemiptera, bugs); orig ...
in the 1980s, requiring the replanting of most of Napa and Sonoma, with disastrous financial consequences. Those who resisted the urge to use AxR-1, such as David Bennion of Ridge Vineyards, saw their vineyards spared from phylloxera damage.


Apple rootstock

Apple rootstocks are used for apple trees and are often the deciding factor of the size of the tree that is grafted onto the root. Dwarfing, semi-dwarf, semi-standard and standard are the size benchmarks for the different sizes of roots that will be grown, with the standard being the largest and dwarf being the smallest. Much of the world's apple production is now using dwarf rootstocks to improve efficiency, increase density and increase yields of fruit per acre. The following is a list of the dwarfing rootstock that are commonly used today in apple production:


Malling rootstocks

Malling 7 rootstock is slightly bigger than an M26 rootstock, and also requires staking in the first several years to establish a centre leader. This rootstock is moderately susceptible to blight and collar rot. The fruit size that M7 produces is good but not as large as an M26, or M9 rootstock. This root is highly susceptible to suckering and leaning over in its later years of life, which is very annoying and causes issues for the producer. Malling 9 rootstock is the most common and well known dwarfing rootstock. This rootstock should be planted in a well-drained site, and requires staking for the duration of its life. This rootstock is also very susceptible to fireblight and burr knots. There have been many clones made of this rootstock, including M.9 NAKB 337, M.9EMLA and M.9. Pajam. Malling 26 rootstock will grow a larger tree than the M9 rootstock will, and is about 40-50% the size of a standard tree. This root is considered very productive and early bearing, and requires staking in the first few years of its life.


Malling Merton rootstocks

Malling-Merton 106 rootstock is slightly smaller than MM 111, but is a very productive tree and has early fruiting abilities. It is a great rootstock to be used in a variety of soil conditions because it is very hardy with moderate vigour. This rootstock must be planted in well-drained soils as it is susceptible to collar rot. Malling-Merton 111 rootstock is one of the biggest and vigorous rootstocks that is used today in commercial orchards, and is about 80-90% the size of a standard-sized tree. It is generally quite winter hardy and produces few burr knots and root suckers. This rootstock is much less popular than it once was, because many commercial producers are now planting higher density orchards, which M111 is not conducive to.


Tree Size


Other rootstocks

R 5 = Robusta 5, O 3 = Ottawa 3, M 27 = Malling 27, a) size compared to seedling of French crab.


See also

* Propagation of grapevines * Fruit tree propagation *
Malling series The Malling series is a group of rootstocks for grafting apple trees. It was developed at the East Malling Research Station of the South-Eastern Agricultural College at Wye in Kent, England. From about 1912, Ronald Hatton and his colleagues ...
** Apple rootstocks ** Cherry rootstocks ** Pear rootstocks ** Plum rootstocks ** Citrus rootstock


Notes


External links

* * {{Viticulture Horticulture Plant reproduction Plant roots