Riocerida, originally named as Rioceratida, is an extinct order of

Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era, and the second of twelve periods of the Phanerozoic Eon (geology), Eon. The Ordovician spans 41.6 million years f ...

nautiloid Nautiloids are a group of cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and species rich, with over 2,500 recorded species. Th ...

cephalopods A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...

. They were the earliest-diverging members of the major group

Orthoceratoidea Orthoceratoidea, from Ancient Greek ὀρθός (''orthós''), meaning "straight", and κέρας (''kéras''), meaning "horn", is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly ...

, with which they share dorsomyarian muscle scars (a few large muscle scars concentrated at the top of the

body chamber The body whorl is part of the morphology of the shell in those gastropod mollusks that possess a coiled shell. The term is also sometimes used in a similar way to describe the shell of a cephalopod mollusk. In gastropods In gastropods, the b ...

). However, they also possess several

plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, an ...

("primitive") traits which are absent in other orthoceratoids:

Cameral Cameralism ( German: ''Kameralismus'') was a German school of public finance, administration and economic management in the 18th and early 19th centuries that aimed at strong management of a centralized economy for the benefit mainly of the st ...

and endosiphuncular deposits are absent, and the

siphuncle The siphuncle is a strand of biological tissue, tissue passing longitudinally through the mollusc shell, shell of a cephalopod mollusc. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the li ...

has a marginal position along the lower edge of the shell. The conch (external shell) shape is slender, usually

orthoconic An orthocone is the long, cone-shaped shell belonging to several species of ancient nautiloid cephalopod—the prehistoric ancestors of today's marine cephalopod mollusks, including the cuttlefishes, nautiluses, octopuses and squids.; During the ...

(straight) or weakly cyrtoconic (slightly curved). The

monophyly In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent comm ...

of Riocerida as a unique group is plausible but difficult to confirm, and they may instead be a

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

grade Grade most commonly refers to: * Grading in education, a measurement of a student's performance by educational assessment (e.g. A, pass, etc.) * A designation for students, classes and curricula indicating the number of the year a student has reach ...

polyphyletic A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...

assemblage ancestral to all later orthoceratoids.

References

{{Taxonbar, from=Q112577892 Prehistoric nautiloids Prehistoric cephalopod orders Orthoceratoidea Early Ordovician first appearances Late Ordovician extinctions