The rinkhals (; ''Hemachatus haemachatus''), also known as the ringhals or ring-necked spitting cobra, is a
species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
venomous snake
''Venomous snakes'' are species of the suborder Serpentes that are capable of producing venom, which they use for killing prey, for defense, and to assist with digestion of their prey. The venom is typically delivered by injection using hollow ...
in the family
Elapidae
Elapidae (, commonly known as elapids , from , variant of "sea-fish") is a family of snakes characterized by their permanently erect fangs at the front of the mouth. Most elapids are venomous, with the exception of the genus '' Emydocephalus ...
. The species is found in parts of southern Africa.
[ It is not a true ]cobra
COBRA or Cobra, often stylized as CoBrA, was a European avant-garde art group active from 1948 to 1951. The name was coined in 1948 by Christian Dotremont from the initials of the members' home countries' capital cities: Copenhagen (Co), Brussels ...
in that it does not belong to the genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Naja
''Naja'' is a genus of venomous elapid snakes commonly known as cobras (or "true cobras"). Various species occur throughout Africa, Southwest Asia, South Asia, and Southeast Asia. Several other elapid species are often called "cobras", such a ...
'', but instead belongs to the monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus ''Hemachatus''. While rinkhals bear a great resemblance to true cobras, they also possess some remarkable differences from these, resulting in their placement outside the genus ''Naja''.
In 2023, the Zimbabwe population was described as a new species, ''H. nyangensis''.
Description
Colouration of the rinkhals varies throughout its distribution area, but a characteristic of the species is that the belly is dark with one or two light-coloured crossbands on the throat. Its usual total length (tail included) is . Some individuals may have a mostly black body, while others are striped. The dorsal scales
In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
are distinct from those of ''Naja'' cobras in that they are keeled. Also unlike members of the genus ''Naja
''Naja'' is a genus of venomous elapid snakes commonly known as cobras (or "true cobras"). Various species occur throughout Africa, Southwest Asia, South Asia, and Southeast Asia. Several other elapid species are often called "cobras", such a ...
'', the rinkhals lacks solid teeth on the maxilla.
Scale pattern
Scalation:
*dorsal scales are keeled[ Branch, Bill (2004). ''Field Guide to Snakes and other Reptiles of Southern Africa''. Third Revised edition, Second impression. Sanibel Island, Florida: Ralph Curtis Books. 399 pp. .]
*17–19 rows of dorsal scales
In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
at midbody
*116–150 ventral scales
In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
* anal plate is entire
*30–47 subcaudal scales
In snakes, the subcaudal scales are the enlarged plates on the underside of the tail. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Comstock Publishing Associates, a Division of Cornell University Press. (7t ...
, paired
*7 upper labial scales
The labial scales are the scales of snakes and other scaled reptiles that border the mouth opening. These do not include the median scales on the upper and lower jawsWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7 ...
*upper labials 3 and 4 entering the eye
*1 preocular
An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system.
In higher organisms, the eye ...
(rarely up to 3)
*3 postoculars
*8–9 lower labials
Geographic range
This species, ''Hemachatus haemachatus'', is found in the Western and Eastern Cape provinces of South Africa
South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
, northeast through the Free State, Lesotho
Lesotho, formally the Kingdom of Lesotho and formerly known as Basutoland, is a landlocked country in Southern Africa. Entirely surrounded by South Africa, it is the largest of only three sovereign enclave and exclave, enclaves in the world, t ...
, KwaZulu-Natal
KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
, South Africa, Western Eswatini
Eswatini, formally the Kingdom of Eswatini, also known by its former official names Swaziland and the Kingdom of Swaziland, is a landlocked country in Southern Africa. It is bordered by South Africa on all sides except the northeast, where i ...
, Mpumalanga
Mpumalanga () is one of the nine provinces of South Africa. The name means "East", or literally "The Place Where the Sun Rises" in the Nguni languages. Mpumalanga lies in eastern South Africa, bordering Eswatini and Mozambique. It shares bor ...
and parts of Gauteng
Gauteng ( , ; Sotho-Tswana languages, Sotho-Tswana for 'place of gold'; or ) is one of the nine provinces of South Africa.
Situated on the Highveld, Gauteng is the smallest province by land area in South Africa. Although Gauteng accounts f ...
, South Africa. An isolated population is centered on Inyanga on the Zimbabwe
file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map
Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
-Mozambique
Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...
border. There are no recent records from this population; so scientists assume this population may be extinct.[
]
Habitat
The rinkhals generally prefers grassland
A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
habitats ranging from the coast to altitudes of up to . The rinkhals is extremely common on the Highveld
The Highveld (Afrikaans: ''Hoëveld,'' , ) is the portion of the South African inland plateau which has an altitude above roughly , but below , thus excluding the Lesotho mountain regions to the south-east of the Highveld. It is home to some of t ...
and in the grasslands of KwaZulu-Natal
KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
. Since the rinkhals has a natural propensity to eat frogs and toads, it can also commonly be found near wetlands in its range.
Behaviour and diet
The rinkhals is diurnal, and hunts during the daytime hours. The rinkhals has a varied diet, and will prey upon any vertebrate that is small enough for it to eat. The rinkhals will most commonly eat toads
Toad (also known as a hoptoad) is a common name for certain frogs, especially of the family Bufonidae, that are characterized by dry, leathery skin, short legs, and large bumps covering the parotoid glands.
In popular culture (folk taxonomy), ...
in the wild, but it has also been recorded to consume rodents and other small vertebrates such as lizards.
When distressed the rinkhals will spread its hood, showing its distinctive striped neck. As a defense mechanism, the rinkhals will spit venom. It will normally spit from a reared position, lunging and hissing while doing so. The rinkhals has ample control over its venom glands, and can accurately spit venom at ranges up to . The rinkhals is also known to fake death, a behavior seen in other snakes such as those of the genus ''Heterodon
''Heterodon'' is a genus of harmless colubrid snakes endemic to North America.Platt, Dwight R. 1983. Heterodon'. Catalogue of American Amphibians and Reptiles. Society for the Study of Amphibians and Reptiles, 351: 1-2. They are stout with upt ...
''. A rinkhals engaging in thanatosis will roll over onto its back, open its mouth, and stick out its toungue.
If all other methods of defense fail, the rinkhals will bite to ward off predators. Since it relies on other forms of defense, bites are usually very rare. Bites most often occur when people try to handle the snake.
Reproduction
Unlike members of the genus ''Naja
''Naja'' is a genus of venomous elapid snakes commonly known as cobras (or "true cobras"). Various species occur throughout Africa, Southwest Asia, South Asia, and Southeast Asia. Several other elapid species are often called "cobras", such a ...
'', the rinkhals is ovoviviparous
Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a "bridging" form of reproduction between egg-laying oviparity, oviparous and live-bearing viviparity, viviparous reproduction. Ovoviviparous animals possess embryos that develo ...
. Instead of laying eggs, the rinkhals gives birth to live young. Litter
Litter consists of waste products that have been discarded incorrectly, without consent, at an unsuitable location. The waste is objects, often man-made, such as aluminum cans, paper cups, food wrappers, cardboard boxes or plastic bottles, but ...
size is usually around twenty to thirty live young, but some females can give birth to over sixty live young. The rinkhals usually gives birth in the late summer, which in the southern hemisphere corresponds to the months ranging from December to March.
Venom
The venom
Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
of the rinkhals is neurotoxic
Neurotoxicity is a form of toxicity in which a biological, chemical, or physical agent produces an adverse effect on the structure or function of the central and/or peripheral nervous system. It occurs when exposure to a substance – specifical ...
and partially cytotoxic
Cytotoxicity is the quality of being toxic to cells. Examples of toxic agents are toxic metals, toxic chemicals, microbe neurotoxins, radiation particles and even specific neurotransmitters when the system is out of balance. Also some types of dr ...
, and is less viscous
Viscosity is a measure of a fluid's rate-dependent resistance to a change in shape or to movement of its neighboring portions relative to one another. For liquids, it corresponds to the informal concept of ''thickness''; for example, syrup h ...
than that of other African elapids
Elapidae (, commonly known as elapids , from , variant of "sea-fish") is a family of snakes characterized by their permanently erect fangs at the front of the mouth. Most elapids are venomous, with the exception of the genus '' Emydocephalus' ...
. When confronting a human, it generally aims its venom at the face. If the venom gets injected, it causes a great amount of pain and even necrosis
Necrosis () is a form of cell injury which results in the premature death of cells in living tissue by autolysis. The term "necrosis" came about in the mid-19th century and is commonly attributed to German pathologist Rudolf Virchow, who i ...
due to the cytotoxic effect. Local symptoms of swelling and bruising are reported in about 25% (a quarter) of cases. General symptoms of drowsiness, nausea
Nausea is a diffuse sensation of unease and discomfort, sometimes perceived as an urge to vomit. It can be a debilitating symptom if prolonged and has been described as placing discomfort on the chest, abdomen, or back of the throat.
Over 30 d ...
, vomiting, violent abdominal pain, cramps and vertigo
Vertigo is a condition in which a person has the sensation that they are moving, or that objects around them are moving, when they are not. Often it feels like a spinning or swaying movement. It may be associated with nausea, vomiting, perspira ...
often occur, as does a mild pyrexia
Fever or pyrexia in humans is a symptom of an anti-infection defense mechanism that appears with body temperature exceeding the normal range caused by an increase in the body's temperature set point in the hypothalamus. There is no single agre ...
l reaction. Polyvalent antivenom is effective for the treatment of rinkhals envenomation, and a fatality from a rinkhals bite has not been recorded in over 40 years.
If the venom enters the eyes, it causes great pain and inflammation. Corneal damage can occur if left untreated, although timely first aid and eye-washing will usually lead to recovery after a few days.
A polyvalent antivenom exists in South Africa and is manufactured by the South African Vaccine Producers.A polyvalent antivenom is currently being developed by the Universidad de Costa Rica
The University of Costa Rica (Spanish: ''Universidad de Costa Rica,'' abbreviated UCR) is a public university in the Republic of Costa Rica, in Central America. Its main campus, Ciudad Universitaria Rodrigo Facio, is located in San Pedro, Co ...
's Instituto Clodomiro Picado.
References
Further reading
*
External links
{{Taxonbar, from=Q169936, from2=Q15148378
Elapidae
Snakes of Africa
Reptiles of Eswatini
Reptiles of Lesotho
Reptiles of South Africa
Reptiles of Zimbabwe
Reptiles described in 1790
Taxa named by Pierre Joseph Bonnaterre
Afrikaans words and phrases