''Rhamphastosula'' ("toucan

booby A booby is a seabird in the genus ''Sula'', part of the family Sulidae. Boobies are closely related to the gannets (''Morus''), which were formerly included in ''Sula''. Systematics and evolution The genus ''Sula'' was introduced by the Fre ...

") is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of sulid bird known from

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s discovered in early

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * ...

and only named species is ''R. ramirezi'', named for Peruvian vertebrate paleontologist Gregorio Ramirez. The genus name is a combination of "Rhamphastos", "toucan" and "sula", an Icelandic word for "fool" that has been used to describe boobies (Sulidae, '' Sula'') in general. ''Rhamphastosula'' is known from a partial skull that has bony characters placing it in the booby family Sulidae. Unlike living boobies, it has a large bill with a convex upper mandible, reminiscent of a toucan. The remains were recovered from the west side of Sud Sacaco Level (4–5 Million year-old) of the Pisco Formation. This area was a sheltered beach with rock platforms.

References

Sulidae Pliocene birds Pliocene Peru Pliocene birds of South America Fossil taxa described in 2004 {{Suliformes-stub