''Quercus manzanillana'' is a shrub in the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...

'' indigenous to

Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...

. It was described in 1924 by

William Trelease William Trelease (February 22, 1857 – January 1, 1945) was an American botanist, entomologist, explorer, writer and educator. This botanist is denoted by the author abbreviation Trel. when citing a botanical name. Trelease was born in Moun ...

. It is found in the Mexican state of

Puebla Puebla, officially the Free and Sovereign State of Puebla, is one of the 31 states that, along with Mexico City, comprise the Federal Entities of Mexico. It is divided into 217 municipalities and its capital is Puebla City. Part of east-centr ...

Description

The twigs of ''Quercus manzanillana'' measure about 2-3 mm in diameter. They are fluted and initially covered in a grayish, tomentulose coating. The

buds In botany, a bud is an undeveloped or Plant embryogenesis, embryonic Shoot (botany), shoot and normally occurs in the axil of a leaf or at the tip of a Plant stem, stem. Once formed, a bud may remain for some time in a dormancy, dormant conditi ...

are light brown,

glabrous Glabrousness () is the technical term for a lack of hair, down, setae, trichomes, or other such covering. A glabrous surface may be a natural characteristic of all or part of a plant or animal, or be due to loss because of a physical condition, ...

and glossy,

ovoid An oval () is a closed curve in a plane which resembles the outline of an egg. The term is not very specific, but in some areas of mathematics (projective geometry, technical drawing, etc.), it is given a more precise definition, which may inc ...

in shape, and approximately 2 x 3 mm in size. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

of this oak species are

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

and elliptical in shape. They are obtuse and exhibit a more or less

cordate Cordate is an adjective meaning ' heart-shaped' and is most typically used for: * Cordate (leaf shape), in plants * Cordate axe, a prehistoric stone tool See also * Chordate A chordate ( ) is a bilaterian animal belonging to the phyl ...

base. The leaf margins are typically entire or very obscurely repand towards the upper part. The leaves measure 2.5-3.5 cm in width and 5-8 cm in length. The upper leaf surface is essentially glabrous and somewhat glossy, while the lower surface is densely covered in a persistent rusty-tomentose pubescence. The leaves have approximately 10 pairs of looped veins, and the

petiole Petiole may refer to: *Petiole (botany), the stalk of a leaf, attaching the blade to the stem *Petiole (insect anatomy) In entomology, petiole is the technical term for the narrow waist of some hymenopteran insects, especially ants, bees, and ...

is tomentulose and measures around 5 mm in length.Trel. (1924), In: Mem. Natl. Acad. Sci. 20: 80

Validity

According to Susana Valencia-A. in 2004, this

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

described by Trelease is not valid. They claim that ''Quercus manzanillana'' represents a

hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...

between different oak species or a morphological

variety Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...

of an existing species. In 2003, Zavala-Chavez proposed an alternative classification for ''Quercus manzanillana''. According to Zavala-Chavez's research, this oak species could be considered a variety of '' Quercus deserticola''. However, these proposals are not universally accepted, and the taxonomic relationship between ''Quercus manzanillana'' and other oak species requires additional investigation.

References

{{Taxonbar, from=Q15338355 manzanillana Flora of Puebla