The principle of faunal succession, also known as the law of faunal succession, is based on the observation that sedimentary rock
strata
In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as e ...
contain
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
ized
flora
Flora is all the plant life present in a particular region or time, generally the naturally occurring ( indigenous) native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' gut flora'' or '' skin flora''.
...
and
fauna
Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is '' flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. ...
, and that these fossils succeed each other vertically in a specific, reliable order that can be identified over wide horizontal distances. A fossilized
Neanderthal
Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an Extinction, extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ag ...
bone will never be found in the same stratum as a fossilized
Megalosaurus, for example, because neanderthals and megalosaurs lived during different
geological periods, separated by many millions of years. This allows for strata to be identified and dated by the fossils found within.
This principle, which received its name from the English
geologist
A geologist is a scientist who studies the solid, liquid, and gaseous matter that constitutes Earth and other terrestrial planets, as well as the processes that shape them. Geologists usually study geology, earth science, or geophysics, althou ...
William Smith, is of great importance in determining the relative age of rocks and strata. The fossil content of rocks together with the
law of superposition helps to determine the time sequence in which sedimentary rocks were laid down.
Evolution
Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
explains the observed faunal and floral succession preserved in rocks. Faunal succession was documented by Smith in England during the first decade of the 19th century, and concurrently in France by
Cuvier
Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in nat ...
(with the assistance of the mineralogist
Alexandre Brongniart
Alexandre Brongniart (5 February 17707 October 1847) was a French chemist, mineralogist, geologist, paleontologist, and zoologist, who collaborated with Georges Cuvier on a study of the geology of the region around Paris. Observing fossil content ...
). Archaic biological features and organisms are succeeded in the
fossil record by more modern versions. For instance,
paleontologists investigating the evolution of
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s predicted that
feather
Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premie ...
s would first be seen in primitive forms on flightless predecessor organisms such as
feathered dinosaur
A feathered dinosaur is any species of dinosaur possessing feathers. While this includes all species of birds, there is a hypothesis that many, if not all non-avian dinosaur species also possessed feathers in some shape or form.
It has been s ...
s. This is precisely what has been discovered in the fossil record: simple feathers, incapable of supporting flight, are succeeded by increasingly large and complex feathers.
In practice, the most useful diagnostic species are those with the fastest rate of species turnover and the widest distribution; their study is termed
biostratigraphy
Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of ...
, the science of dating rocks by using the
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s contained within them. In
Cenozoic
The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configu ...
strata, fossilized tests of
foraminifera
Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly ...
are often used to determine faunal succession on a refined scale, each biostratigraphic unit (
biozone) being a geological stratum that is defined on the basis of its characteristic fossil
taxa. An outline microfaunal zonal scheme based on both foraminifera and
ostracod
Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typic ...
a was compiled by M. B. Hart (1972).
Earlier fossil life forms are simpler than more recent forms, and more recent fossil forms are more similar to living forms (principle of faunal succession).
[''Evolutionary Analysis'', 4th Edition. p 61.]
See also
*
Law of superposition
*
Principle of cross-cutting relationships
*
Principle of lateral continuity
*
Principle of original horizontality
References
{{Geologic Principles
Paleontological concepts and hypotheses
Dating methods
Stratigraphy
de:Biostratigraphie#Das Prinzip der Fossilfolge