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''Populus tremula'' (commonly called aspen, common aspen, Eurasian aspen, European aspen, or quaking aspen) is a species of poplar native to cool
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
regions of Europe and Asia, from Iceland and the British IslesJames Kilkell
Irish native Aspen tree
/ref> east to
Kamchatka The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and west ...
, north to inside the
Arctic Circle The Arctic Circle is one of the two polar circles, and the most northerly of the five major circles of latitude as shown on maps of Earth. Its southern equivalent is the Antarctic Circle. The Arctic Circle marks the southernmost latitude at ...
in Scandinavia and northern Russia, and south to central Spain, Turkey, the
Tian Shan The Tian Shan,, , otk, 𐰴𐰣 𐱅𐰭𐰼𐰃, , tr, Tanrı Dağı, mn, Тэнгэр уул, , ug, تەڭرىتاغ, , , kk, Тәңіртауы / Алатау, , , ky, Теңир-Тоо / Ала-Тоо, , , uz, Tyan-Shan / Tangritog‘ ...
, North Korea, and northern Japan. It also occurs at one site in northwest Africa in Algeria. In the south of its range, it occurs at high altitudes in mountains.Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .Den Virtuella Floran
''Populus tremula'' (in Swedish; with maps)
/ref>


Description

It is a substantial
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing to tall by broad, with a trunk attaining over in diameter. The
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, e ...
is pale greenish-grey and smooth on young trees with dark grey diamond-shaped
lenticel A lenticel is a porous tissue consisting of cells with large intercellular spaces in the periderm of the secondarily thickened organs and the bark of woody stems and roots of dicotyledonous flowering plants. It functions as a pore, providing a ...
s, becoming dark grey and fissured on older trees. The adult
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
, produced on branches of mature trees, are nearly round, slightly wider than long, diameter, with a coarsely toothed margin and a laterally flattened petiole long. The flat petiole allows them to tremble in even slight breezes, and is the source of its scientific name, as well as one of its vernacular names "langues de femmes" attributed to Gerard's 17th-century ''Herball''. The leaves on seedlings and fast-growing stems of suckers (root sprouts) are of a different shape, heart-shaped to nearly triangular. They are also often much larger, up to long; their petiole is also less flattened. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s are wind-pollinated
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in '' Salix''). They contain many, usually unisexual flowers, arrang ...
s produced in early spring before the new leaves appear; they are
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
, with male and female catkins on different trees. The male catkins are patterned green and brown, long when shedding pollen; the female catkins are green, long at pollination, maturing in early summer to bear 10–20 (50–80) capsules each containing numerous tiny
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s embedded in downy fluff. The fluff assists wind dispersal of the seeds when the capsules split open at maturity.Trees for Life Species Profile
Aspen
It can be distinguished from the closely related North American ''
Populus tremuloides ''Populus tremuloides'' is a deciduous tree native to cooler areas of North America, one of several species referred to by the common name aspen. It is commonly called quaking aspen, trembling aspen, American aspen, mountain or golden aspen, tr ...
'' by the leaves being more coarsely toothed. Like other aspens, it spreads extensively by suckers (root sprouts), which may be produced up to 40 m from the parent tree, forming extensive clonal colonies. This often makes the job of clearing unwanted trees from an area especially difficult, as new suckers will continue to sprout from the extensive root system for up to several years after all surface growth has been eliminated.


Ecology

Eurasian aspen is a water and light demanding species that is able to vigorously colonize an open area after fire, clear cutting or other kind of damages. After an individual has been damaged or destroyed, root suckers are produced abundantly on the shallow lateral roots. Fast growth continues until the age of about 20 years when crown competition increases. After that, growth speed decreases and culminates at about 30 years of age. Aspen can reach an age of 200 years. It is a very hardy species and tolerates long, cold winters and short summers. Aspen is resistant to browsing pressure by
fallow deer ''Dama'' is a genus of deer in the subfamily Cervinae, commonly referred to as fallow deer. Name The name fallow is derived from the deer's pale brown colour. The Latin word ''dāma'' or ''damma'', used for roe deer, gazelles, and antelopes ...
due to its unpleasant taste. This species is important for the hornet moth, which uses it as a host during the larval stage.


Fossil record

Fossils of ''Populus tremula'' have been described from the fossil flora of Kızılcahamam district in
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
which is of early
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58fastigiate
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
‘Erecta’, with bright yellow autumn colouring, has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
’s
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
. The cultivar is colloquially known as "Swedish columnar" in Canada and the United States. The hybrid with ''
Populus alba ''Populus alba'', commonly called silver poplar,Webb, C. J.; Sykes, W. R.; Garnock-Jones, P. J. 1988: Flora of New Zealand. Vol. IV. Naturalised Pteridophytes, Gymnosperms, Dicotyledons. 4. Christchurch, New Zealand, Botany Division, D.S.I.R. si ...
'' (white poplar), known as grey poplar, ''Populus'' × ''canescens'', is widely found in Europe and central Asia. Hybrids with several other aspens have also been bred at forestry research institutes in order to find trees with greater timber production and disease resistance (e.g. ''P. tremula'' × ''P. tremuloides'', bred in DenmarkJensen, N. (1994). Guide til Arboretet i Hørsholm (in Danish).).


Use

The wood of aspen is light and soft with very little shrinkage. It is used for
lumber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
and
matches A match is a tool for starting a fire. Typically, matches are made of small wooden sticks or stiff paper. One end is coated with a material that can be ignited by friction generated by striking the match against a suitable surface. Wooden matc ...
but is also valued in the
pulp and paper industry The pulp and paper industry comprises companies that use wood as raw material and produce Pulp (paper), pulp, paper, paperboard and other cellulose-based products. Manufacturing process The pulp is fed to a paper machine where it is formed as ...
, being particularly useful for writing paper. In addition, it is used for
plywood Plywood is a material manufactured from thin layers or "plies" of wood veneer that are glued together with adjacent layers having their wood grain rotated up to 90 degrees to one another. It is an engineered wood from the family of manufactured ...
and different types of flake and particle boards. Given its hardiness and capacity for rapid growth and regeneration, it plays an important role in the production of wood for
renewable energy Renewable energy is energy that is collected from renewable resources that are naturally replenished on a human timescale. It includes sources such as sunlight, wind, the movement of water, and geothermal heat. Although most renewable energy ...
. Ecologically, the species is important as many insect and fungi species benefit from it. The tree further provides
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
for several mammals and birds that require young forests.


References


External links


Video, commentary and annotation on why Aspen leaves tremble.

''Populus tremula''
- distribution map, genetic conservation units and related resources.
European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that supports the conservation and sustainable use of forest genetic resources in Europe. The programme’s tasks include to coordinate and promote '' in situ'' an ...
(EUFORGEN) *
Populus tremula
' ''-'' Aspen; info, images and video a
Woodland Trust
* {{Authority control tremula Flora of temperate Asia Flora of Europe Flora of Algeria Trees of continental subarctic climate Trees of humid continental climate Trees of subpolar oceanic climate Plants described in 1753 Taxa named by Carl Linnaeus