Polyphaga is the largest and most diverse suborder of

beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...

s. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the beetle species so far discovered. Key characteristics of Polyphaga are that the hind coxa (base of the leg) does not divide the first and second abdominal/ventral plates which are known as sternites. Also, the

notopleural The notopleuron (plural notopleura) is a region on an insect thorax. Notopleura are useful in characterizing species, particularly, though not uniquely, in the Order Diptera (the "true flies"). The notopleuron is a thoracic pleurite (a sclerite ...

suture (found under the pronotal shield) is not present.

Etymology

The name of ''polyphaga'' is derived from two Greek words: , meaning 'many', and , meaning 'to eat', so the suborder is called the “eaters of many things”.

Classification

The five main infraorders are: * Bostrichiformia — including furniture beetles and skin beetles * Cucujiformia — includes

lady beetle Coccinellidae () is a widespread family of small beetles ranging in size from . They are commonly known as ladybugs in North America and ladybirds in Great Britain. Some entomologists prefer the names ladybird beetles or lady beetles as they ...

longhorn beetle The longhorn beetles (Cerambycidae), also known as long-horned or longicorns, are a large family of beetles, with over 35,000 species described. Most species are characterized by extremely long antennae, which are often as long as or longer than ...

s, weevils,

checkered beetle Cleridae are a family of beetles of the superfamily Cleroidea. They are commonly known as checkered beetles. The family Cleridae has a worldwide distribution, and a variety of habitats and feeding preferences. Cleridae have many niches and f ...

s and leaf beetles * Elateriformia — includes click beetles and fireflies * Scarabaeiformia — includes scarab beetles, stag beetles, and dung beetles *

Staphyliniformia Staphyliniformia is a large infraorder of beetles. It contains over 70,000 described species from all regions of the world. Most species occur in moist habitats - various kinds of rotting plant debris, fungi, dung, carrion, many live in fresh wat ...

— includes rove beetles and water scavenger beetles Phylogenetic studies have also suggested that Scirtoidea ( Scirtidae, Decliniidae), Clamboidea ( Clambidae, Derodontidae, Eucinetidae), ''

Rhinorhipus ''Rhinorhipus'' is a genus of beetles that contains a single species, ''Rhinorhipus tamborinensis'' from southern Queensland, Australia. It is the sole member of the family Rhinorhipidae and superfamily Rhinorhipoidea. It is an isolated lineage ...

'' and Nosodendridae are independent lineages of Polyphaga that lie outside these groups. The internal classification of Polyphaga involves several superfamilies or series, whose constituents are relatively stable, although some smaller families (whose rank even is disputed) are allocated to different clades by different authors. Large superfamilies include Hydrophiloidea, Staphylinoidea, Scarabaeoidea,

Buprestoidea Buprestoidea is a superfamily of beetles. It contains two families: * Buprestidae Leach 1815, the jewel beetle Buprestidae is a family of beetles known as jewel beetles or metallic wood-boring beetles because of their glossy iridescent colo ...

, Byrrhoidea, Elateroidea, and

Bostrichoidea Bostrichoidea is a superfamily of beetles. It is the type superfamily of the infraorder Bostrichiformia. Description The Bostrichoidea are united by the following features: modified cryptonephridism, the structure of the aedeagus in males, and ...

. The infraorder Cucujiformia includes the vast majority of phytophagous (plant-eating) beetles, united by cryptonephric Malpighian tubules of the normal type, a cone ommatidium with open rhabdom, and lack of functional spiracles on the eighth abdominal segment. Constituent superfamilies of Cucujiformia are Cleroidea, Cucujoidea, Tenebrionoidea, Chrysomeloidea, and Curculionoidea. Evidently adoption of a phytophagous lifestyle correlates with taxon diversity in beetles, with Cucujiformia, especially weevils ( Curculionoidea), forming a major radiation.

References

External links

Phylogeny of Insects

Site devoted to Staphyliniformia

site devoted to stag beetles

news article highlighting the damage caused by Longhorn beetles
* Insect suborders {{Polyphaga-stub

Etymology

Classification

See also

References

External links