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''Pinus muricata'', the bishop pine, is a
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanic ...
with a very restricted range: mostly in California, including several offshore
Channel Islands The Channel Islands ( nrf, Îles d'la Manche; french: îles Anglo-Normandes or ''îles de la Manche'') are an archipelago in the English Channel, off the French coast of Normandy. They include two Crown Dependencies: the Bailiwick of Jersey, ...
, and a few locations in Baja California, Mexico. It is always on or near the coast. In San Luis Obispo County it is found alone or in stands scattered on the coastal mountains and hills from
Morro Bay Morro Bay (''Morro'', Spanish for "Hill") is a seaside city in San Luis Obispo County, California. Located on the Central Coast of California, the city population was 10,757 as of the 2020 census, up from 10,234 at the 2010 census. The town ...
to Shell Beach. A few stands of the tree are seen on the hills above the Sycamore Canyon Resort in
Avila Beach Avila Beach (Spanish: ''Ávila'') is an unincorporated community in San Luis Obispo County, California, United States, located on San Luis Obispo Bay about 160 miles (257 km) northwest of Los Angeles, and about south of San Francisco. The ...
. Within the City of
San Luis Obispo San Luis Obispo (; Spanish for " St. Louis the Bishop", ; Chumash: ''tiłhini'') is a city and county seat of San Luis Obispo County, in the U.S. state of California. Located on the Central Coast of California, San Luis Obispo is roughly hal ...
, the Terrace Hill Open Space has several scattered specimens. Bishop pine seems to prefer already disturbed, unvegetated areas where it probably faces less competition from oaks and shrubs. The common name "bishop pine" resulted from the tree having been first identified near the Mission of San Luis Obispo in
San Luis Obispo San Luis Obispo (; Spanish for " St. Louis the Bishop", ; Chumash: ''tiłhini'') is a city and county seat of San Luis Obispo County, in the U.S. state of California. Located on the Central Coast of California, San Luis Obispo is roughly hal ...
, California. This tree has a large number of common names and other prior scientific names, due primarily to numerous variant forms. Other English names that have occasionally been used are prickle cone pine, Obispo pine, Santa Cruz pine and dwarf marine pine.


Description

''Pinus muricata'' is a
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ext ...
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing to a height of , rarely up to , with a trunk diameter of up to . The
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
is often smaller, stunted and twisted in coastal exposures. It is drought-tolerant and grows on dry, rocky soil. The needles are in pairs, green to blue-green, and long.
Cone A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines con ...
s occur in one to five clusters. The cones are strongly reflexed down the branch, long; the scales are stiff, thin on the side of the cone facing the stem, but greatly thickened on the side facing away and with a stout spine; both features adaptive to minimise
squirrel Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrels. ...
predation and fire damage to the cones. The cones remain unopened for many years until fire or strong heat causes them to open and release the seeds. File:Pinus muricata-1.jpg, Needles File:Pinus muricata, Morro Bay State Park, San Luis Obispo, California 4.jpg, Cones File:Growing tip of Pinus muricata.JPG, Growing tips


Forms

There are two ''Pinus muricata'' forms: * a southern form with bright green needles * a northern form with dark blue-green needles. The
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on nat ...
composition also differs. The dividing line between the two is very sharp, five miles (8 km) south of the boundary between Mendocino County and Sonoma County, California. Experimental attempts to hybridize the two forms have consistently failed, indicating that their taxonomic relationship may be more distant than the very small differences in appearance would suggest.


Ecology

''Pinus muricata'' is found with several oak and cypress associates within the
California Coast Ranges The Coast Ranges of California span from Del Norte or Humboldt County, California, south to Santa Barbara County. The other three coastal California mountain ranges are the Transverse Ranges, Peninsular Ranges and the Klamath Mountains. ...
. There are also a number of common
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abo ...
flora associates including
sword fern Sword fern is a common name for several ferns and may refer to: *''Nephrolepis'', a tropical genus of ferns, especially: **'' Nephrolepis exaltata'', commonly cultivated as a houseplant, including the Boston fern *''Polystichum'', a cosmopolitan ...
,
salal ''Gaultheria shallon'' is an evergreen shrub in the heather family (Ericaceae), native to western North America. In English, it is known as salal, shallon, or (mainly in Britain) gaultheria. Description ''Gaultheria shallon'' is tall, spra ...
and
western poison oak ''Toxicodendron diversilobum'' (syn. ''Rhus diversiloba''), commonly named Pacific poison oak or western poison oak, is a woody vine or shrub in the sumac family, Anacardiaceae. It is widely distributed in western North America, inhabiting coni ...
. Notable occurrences of ''P. muricata'' is in association with Mendocino cypress as a
pygmy forest In anthropology, pygmy peoples are ethnic groups whose average height is unusually short. The term pygmyism is used to describe the phenotype of endemic short stature (as opposed to disproportionate dwarfism occurring in isolated cases in a ...
on coastal terraces in Mendocino County and Sonoma County, including one location within Salt Point State Park. It is classified an
endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
in Mexico.


Uses

''Pinus muricata'' has been used in
plantation A plantation is an agricultural estate, generally centered on a plantation house, meant for farming that specializes in cash crops, usually mainly planted with a single crop, with perhaps ancillary areas for vegetables for eating and so on. Th ...
s with resultant growth rates higher than in the wild, but with adverse impacts to
biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variability''), species ('' species diversity''), and ecosystem ('' ecosystem diversity'') ...
. This plant has ornamental value, and is cultivated in parks and gardens. It has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
.RHS Plant Selector Pinus muricata AGM / RHS Gardening
/ref>


References


External links

* * * * * {{Taxonbar, from=Q866150 muricata Trees of Baja California Trees of the Southwestern United States Flora of California Natural history of the California chaparral and woodlands Trees of mild maritime climate