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Physiological Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemica ...
plant disorders are caused by non-
pathological Pathology is the study of the causes and effects of disease or injury. The word ''pathology'' also refers to the study of disease in general, incorporating a wide range of biology research fields and medical practices. However, when used in th ...
conditions such as poor light, adverse weather, water-logging, phytotoxic compounds or a lack of nutrients, and affect the functioning of the
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae excl ...
system. Physiological disorders are distinguished from plant diseases caused by
pathogen In biology, a pathogen ( el, πάθος, "suffering", "passion" and , "producer of") in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a g ...
s, such as a
virus A virus is a submicroscopic infectious agent that replicates only inside the living cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Since Dmitri Ivanovsk ...
or
fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately fr ...
. While the symptoms of physiological disorders may appear disease-like, they can usually be prevented by altering environmental conditions. However, once a plant shows symptoms of a physiological disorder it is likely that, that season's growth or yield will be reduced.


Diagnosis of disorders

Diagnosis of the cause of a physiological disorder (or disease) can be difficult, but there are many web-based guides that may assist with this. Examples are: ''Abiotic plant disorders: Symptoms, signs and solutions''; ''Georgia Corn Diagnostic Guide''; ''Diagnosing Plant Problems'' (Kentucky); and ''Diagnosing Plant Problems'' (Virginia). Some general tips to diagnosing plant disorders: *Examine where symptoms first appear on a plant—on new
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
, old leaves or all over? *Note the pattern of any discolouration or yellowing—is it all over, between the
vein Veins are blood vessels in humans and most other animals that carry blood towards the heart. Most veins carry deoxygenated blood from the tissues back to the heart; exceptions are the pulmonary and umbilical veins, both of which carry oxygenated ...
s or around the edges? If only the veins are yellow, deficiency is probably not involved. *Note general patterns rather than looking at individual plants—are the symptoms distributed throughout a group of plants of the same type growing together? In the case of a deficiency all of the plants should be similarly effected, although distribution will depend on past treatments applied to the soil. *Soil analysis, such as determining pH, can help to confirm the presence of physiological disorders. *Considering recent conditions, such as heavy rains, dry spells, frosts, etc., may also help to determine the cause of plant disorders.


Weather damage

Frost Frost is a thin layer of ice on a solid surface, which forms from water vapor in an above-freezing atmosphere coming in contact with a solid surface whose temperature is below freezing, and resulting in a phase change from water vapor (a g ...
and cold are major causes of
crop A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydropon ...
damage to tender plants, although hardy plants can also suffer if new growth is exposed to a hard frost following a period of warm weather. Symptoms will often appear overnight, affecting many types of plants. Leaves and stems may turn black, and buds and flowers may be discoloured, and frosted blooms may not produce
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
. Many
annual Annual may refer to: *Annual publication, periodical publications appearing regularly once per year ** Yearbook ** Literary annual *Annual plant *Annual report *Annual giving *Annual, Morocco, a settlement in northeastern Morocco *Annuals (band), ...
plants, or plants grown in frost free areas, can suffer from damage when the air temperature drops below 40 degrees
Fahrenheit The Fahrenheit scale () is a temperature scale based on one proposed in 1724 by the physicist Daniel Gabriel Fahrenheit (1686–1736). It uses the degree Fahrenheit (symbol: °F) as the unit. Several accounts of how he originally defined hi ...
(4 degrees
Celsius The degree Celsius is the unit of temperature on the Celsius scale (originally known as the centigrade scale outside Sweden), one of two temperature scales used in the International System of Units (SI), the other being the Kelvin scale. The d ...
). Tropical plants may begin to experience cold damage when the temperature is 42 to 48 °F (5 to 9 °C), symptoms include wilting of the top of the stems and/or leaves, and blackening or softening of the plant tissue. Frost or cold damage can be avoided by ensuring that tender plants are properly hardened before planting, and that they are not planted too early in the season, before the risk of frost has passed. Avoid planting susceptible plants in frost pockets, or where they will receive early morning sun. Protect young buds and bloom with horticultural fleece if frost is forecast. Cold, drying easterly winds can also severely inhibit spring growth even without an actual frost, thus adequate shelter or the use of windbreaks is important. Drought can cause plants to suffer from water stress and wilt. Adequate
irrigation Irrigation (also referred to as watering) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has been devel ...
is required during prolonged hot, dry periods. Rather than shallow daily watering, during a drought water should be directed towards the roots, ensuring that the soil is thoroughly soaked two or three times a week.
Mulch A mulch is a layer of material applied to the surface of soil. Reasons for applying mulch include conservation of soil moisture, improving fertility and health of the soil, reducing weed growth and enhancing the visual appeal of the area. A mu ...
es also help preserve soil moisture and keep roots cool. Heavy rains, particularly after prolonged dry periods, can also cause roots to split,
onion An onion (''Allium cepa'' L., from Latin ''cepa'' meaning "onion"), also known as the bulb onion or common onion, is a vegetable that is the most widely cultivated species of the genus '' Allium''. The shallot is a botanical variety of the on ...
saddleback (splitting at the base),
tomato The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...
es split and
potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Uni ...
es to become deformed or hollow. Using mulches or adding organic matter such as
leaf mold Leaf mold (spelled leaf mould outside of the United States) is the compost produced by decomposition of shaded deciduous shrub and tree leaves, primarily by fungal breakdown in a slower cooler manner as opposed to the bacterial degradation of leav ...
,
compost Compost is a mixture of ingredients used as plant fertilizer and to improve soil's physical, chemical and biological properties. It is commonly prepared by decomposing plant, food waste, recycling organic materials and manure. The resulting ...
or well rotted
manure Manure is organic matter that is used as organic fertilizer in agriculture. Most manure consists of animal feces; other sources include compost and green manure. Manures contribute to the Soil fertility, fertility of soil by adding organic ma ...
to the
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
will help to act as a 'buffer' between sudden changes in conditions. Water-logging can occur on poorly drained soils, particularly following heavy rains. Plants can become yellow and stunted, and will tend to be more prone to drought and diseases. Improving drainage will help to alleviate this problem. Hail can cause damage to soft skinned fruits, and may also allow brown rot or other
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately fr ...
to penetrate the plant. Brown spot markings or lines on one side of a mature
apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus '' Malus''. The tree originated in Central Asia, where its wild ancest ...
are indicative of a spring hailstorm. Plants affected by
salt Salt is a mineral composed primarily of sodium chloride (NaCl), a chemical compound belonging to the larger class of salts; salt in the form of a natural crystalline mineral is known as rock salt or halite. Salt is present in vast quant ...
stress are able to take water from soil, due to an
osmotic Osmosis (, ) is the spontaneous net movement or diffusion of solvent molecules through a selectively-permeable membrane from a region of high water potential (region of lower solute concentration) to a region of low water potential (region ...
imbalance between soil and plant.


Nutrient deficiencies

Poor growth and a variety of disorders such as leaf discolouration (
chlorosis In botany, chlorosis is a condition in which leaves produce insufficient chlorophyll. As chlorophyll is responsible for the green color of leaves, chlorotic leaves are pale, yellow, or yellow-white. The affected plant has little or no ability to ...
) can be caused by a shortage of one or more plant nutrients. Poor plant uptake of a nutrient from the soil (or other growing medium) may be due to an absolute shortage of that element in the growing medium, or because that element is present in a form that is not available to the plant. The latter can be caused by incorrect pH, shortage of water, poor root growth or an excess of another nutrient. Plant nutrient deficiencies can be avoided or corrected using a variety of approaches including the consultation of experts on-site, the use of
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
and plant-tissue testing services, the application of prescription-blend
fertilizer A fertilizer (American English) or fertiliser (British English; see spelling differences) is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from ...
s, the application of fresh or well-decomposed
organic matter Organic matter, organic material, or natural organic matter refers to the large source of carbon-based compounds found within natural and engineered, terrestrial, and aquatic environments. It is matter composed of organic compounds that have c ...
, and the use of biological systems such as
cover crop In agriculture, cover crops are plants that are planted to cover the soil rather than for the purpose of being harvested. Cover crops manage soil erosion, soil fertility, soil quality, water, weeds, pests, diseases, biodiversity and wildlife in ...
s,
intercropping Intercropping is a multiple cropping practice that involves growing two or more crops in proximity. In other words, intercropping is the cultivation of two or more crops simultaneously on the same field. The most common goal of intercropping is ...
, improved fallows, ley cropping,
permaculture Permaculture is an approach to land management and settlement design that adopts arrangements observed in flourishing natural ecosystems. It includes a set of design principles derived using whole-systems thinking. It applies these principl ...
, or
crop rotation Crop rotation is the practice of growing a series of different types of crops in the same area across a sequence of growing seasons. It reduces reliance on one set of nutrients, pest and weed pressure, and the probability of developing resistant ...
. Nutrient (or mineral) deficiencies include: * Boron deficiency * Calcium deficiency * Iron deficiency * Magnesium deficiency * Manganese deficiency *
Molybdenum deficiency Molybdenum deficiency refers to the clinical consequences of inadequate supplies of molybdenum in the diet. The amount of molybdenum required is relatively small, and molybdenum deficiency usually does not occur in natural settings. However, it ...
*
Nitrogen deficiency Nitrogen deficiency is a deficiency of nitrogen in plants. This can occur when organic matter with high carbon content, such as sawdust, is added to soil. Soil organisms use any nitrogen available to break down carbon sources, making nitrogen una ...
*
Phosphorus deficiency Phosphorus deficiency is a plant disorder associated with insufficient supply of phosphorus. Phosphorus refers here to salts of phosphates (PO43−), monohydrogen phosphate (HPO42−), and dihydrogen phosphate (H2PO4−). These anions readily in ...
* Potassium deficiency * Zinc deficiency


References

{{physiology types 01 Horticulture