Permocupedidae is a family of Protocoleopteran stem group beetles. They first appeared during the Early Permian, and were one of the dominant groups of beetles during the

Middle Permian The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0. ...

. They became rare in the

Late Permian Late or LATE may refer to: Everyday usage * Tardy, or late, not being on time * Late (or the late) may refer to a person who is dead Music * ''Late'' (The 77s album), 2000 * Late (Alvin Batiste album), 1993 * Late!, a pseudonym used by Dave Groh ...

, with only one species known from the

Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...

, '' Frankencupes ultimus'' from the

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is di ...

(

Anisian In the geologic timescale, the Anisian is the lower stage (stratigraphy), stage or earliest geologic age, age of the Middle Triassic series (stratigraphy), series or geologic epoch, epoch and lasted from million years ago until million years ag ...

) Röt Formation of Germany. They are thought to have been xylophagous (wood eating), which is presumed to be the ancestral ecology of beetles.

Taxonomy

Kirejtshuk (2020) included the following genera in an expanded (''

sensu lato ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...

'') family, with the traditional (''

sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...

'') Permocupedidae and related Taldycupedidae as subfamilies Permocupedinae and Taldycupedinae respectively. However, other studies have recovered Talycupedidae as more closely related to crown-group beetles than to Permocupedidae. *'' Afrocupes'' – South Africa:

Western Cape The Western Cape ( ; , ) is a provinces of South Africa, province of South Africa, situated on the south-western coast of the country. It is the List of South African provinces by area, fourth largest of the nine provinces with an area of , an ...

(Kenmoore Farm), Whitehill Formation, Permian, Cisuralian, Sakmarian/Artinskian, −290.1–279.5 Ma. **''Afrocupes firmae'' (Type species) *'' Archicupes'' ( syn ''Palaeocupes'' ) – Russia, Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma **''Archicupes jacobsoni'' (Type species) **''"Archicupes" reichardti'' – considered indeterminate to genus by Kirejtshuk (2020) *'' Cytocupoides'' – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. **''Cytocupoides longatus'' (Type species) *'' Cytocupes'' – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. **''Cytocupes angustus'' (Type species) *'' Eocupes'' – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma). **''Eocupes cellulosus'' **''Eocupes lukjanovitshi'' (Type species) *'' Frankencupes'' – Germany: Gambach am Main (

Lower Franconia Lower Franconia (, ) is one of seven districts of Bavaria, Germany. The districts of Lower, Middle and Upper Franconia make up the region of Franconia. It consists of nine districts and 308 municipalities (including three cities). History After ...

), Röt Formation Middle Triassic, Olenekian/Anisian, −247.2–242.0 Ma. **''Frankencupes ultimus'' (Type species) *'' Ichthyocupes'' – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Kemerovo Region (Tykhta River, Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma. **''Ichthyocupes kuznetskiensis'' **''Ichthyocupes skoki'' **''Ichthyocupes tyzhnovi'' (Type species) *'' Kaltanocoleus'' – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. **''Kaltanocoleus pospelovi'' (Type species) *'' Kaltanicupes'' – Russia: Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Orenburg Region (Kargala), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Sweverodvinian), −265.0–259.0 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. One undescribed species of this genus was recorded from Indian Barakar Formation of the Lower Permian. **''Kaltanicupes acutus'' **''Kaltanicupes kargalensis'' **''Kaltanicupes kitjakensis'' **''Kaltanicupes richteri'' (Type species) *'' Linicupes'' – China:

Anhui Anhui is an inland Provinces of China, province located in East China. Its provincial capital and largest city is Hefei. The province is located across the basins of the Yangtze and Huai rivers, bordering Jiangsu and Zhejiang to the east, Jiang ...

(Houdong, Yinping Mountain, SW Chaohu City), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian/Vyatkian), −265.0–259.0 Ma. **''Linicupes yinpinensis'' (Type species) *'' Maricoleus'' – Russia,

Primorsky Krai Primorsky Krai, informally known as Primorye, is a federal subjects of Russia, federal subject (a krais of Russia, krai) of Russia, part of the Far Eastern Federal District in the Russian Far East. The types of inhabited localities in Russia, ...

(Cape, Novosilsky,

Russky Island Russky Island () is an island in Peter the Great Gulf in the Sea of Japan, in Primorsky Krai, Russia. It is the largest island in the Eugénie Archipelago, separated from the Muravyov-Amursky Peninsula immediately to the north by the Eastern ...

), Lower Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma. **''Maricoleus valentinae'' (Type species) *'' Permocupes'' (syn ''Permocupoides'' ) – Russia: Arkhangel’sk Region (Letopala River and Soyana), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma: Tatarstan (Mendeleevsk District, Tikhie Gory), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. **''Permocupes distinctus'' **''Permocupes grandis'' **''Permocupes latus'' **''Permocupes semenovi'' (Type species) **''Permocupes sojanensis'' *'' Pintolla'' – Brazil:

Rio Grande do Sul Rio Grande do Sul (, ; ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative units of Brazil#List, fifth-most populous state and the List of Brazilian s ...

(Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma. **''Pintolla ponomarenkoi'' (Type species) *'' Protocupoides'' (syn ''Tomiocupes'' , ''Tricupes'' ) – Russia: Vologda Region (Isady locality, Mutovino), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinskian/Vyatkian), −259.0–254.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma; Kemerovo Region (Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. **''Protocupoides acer'' **''Protocupoides carinatus'' **''Protocupoides elongatus'' **''Protocupoides esini'' **''Protocupoides plavilstshikovi'' **''Protocupoides sharovi'' **''"Protocupoides" rohdendorfi'' – Brazil (Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Kungurian, −279.5–272.5 Ma; considered indeterminate to genus by Kirejtshuk (2020) *'' Tatarocupes'' – Russia: Orenburg Region (Novo-Aleksandrovka), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian), −265.0–259.0 Ma. **''Tatarocupes granulatus'' (Type species) *'' Uralocupes'' – Russia: Belebeevo Formation Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. **''Uralocupes major'' (Type species)

References

{{Taxonbar, from=Q19385267 Permocupedoidea Permian insects Permian China Prehistoric animals of China † Prehistoric insect families