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The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
, is a
cosmopolitan Cosmopolitan may refer to: Food and drink * Cosmopolitan (cocktail), also known as a "Cosmo" History * Rootless cosmopolitan, a Soviet derogatory epithet during Joseph Stalin's anti-Semitic campaign of 1949–1953 Hotels and resorts * Cosmopoli ...
bird of prey (
raptor Raptor or RAPTOR may refer to: Animals The word "raptor" refers to several groups of bird-like dinosaurs which primarily capture and subdue/kill prey with their talons. * Raptor (bird) or bird of prey, a bird that primarily hunts and feeds on v ...
) in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Falconidae The falcons and caracaras are around 60 species of diurnal birds of prey that make up the family Falconidae (representing all extant species in the order Falconiformes). The family is divided into three subfamilies, Herpetotherinae, which inclu ...
. A large, crow-sized
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
, it has a blue-grey back, barred white underparts, and a black head. The peregrine is renowned for its speed, reaching over during its characteristic hunting stoop (high-speed dive), making it the fastest bird in the world, as well as the fastest member of the animal kingdom. According to a ''
National Geographic ''National Geographic'' (formerly the ''National Geographic Magazine'', sometimes branded as NAT GEO) is a popular American monthly magazine published by National Geographic Partners. Known for its photojournalism, it is one of the most widel ...
'' TV program, the highest measured speed of a peregrine falcon is . As is typical for bird-eating raptors, peregrine falcons are sexually dimorphic, with females being considerably larger than males. The peregrine's breeding range includes land regions from the Arctic tundra to the
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also refer ...
. It can be found nearly everywhere on Earth, except extreme
polar region The polar regions, also called the frigid zones or polar zones, of Earth are the regions of the planet that surround its geographical poles (the North and South Poles), lying within the polar circles. These high latitudes are dominated by flo ...
s, very high mountains, and most
tropical rainforest Tropical rainforests are rainforests that occur in areas of tropical rainforest climate in which there is no dry season – all months have an average precipitation of at least 60 mm – and may also be referred to as ''lowland equator ...
s; the only major ice-free landmass from which it is entirely absent is
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
. This makes it the world's most widespread raptor, and one of the most widely found bird species. In fact, the only land-based bird species found over a larger geographic area is not always naturally occurring, but one widely introduced by humans, the rock pigeon, which in turn now supports many peregrine populations as a prey species. The peregrine is a highly successful example of
urban wildlife Urban wildlife is wildlife that can live or thrive in urban/suburban environments or around densely populated human settlements such as townships. Some urban wildlife, such as house mice, are synanthropic, ecologically associated with and ev ...
in much of its range, taking advantage of tall buildings as nest sites and an abundance of prey such as pigeons and ducks. Both the English and
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
s of this
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
mean "wandering falcon," referring to the migratory habits of many northern populations. Experts recognize 17 to 19
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
, which vary in appearance and range; disagreement exists over whether the distinctive Barbary falcon is represented by two subspecies of ''Falco peregrinus,'' or is a separate species, ''F. pelegrinoides''. The two species' divergence is relatively recent, during the time of the last ice age, therefore the genetic differential between them (and also the difference in their appearance) is relatively tiny. They are only about 0.6–0.8% genetically differentiated. Although its diet consists almost exclusively of medium-sized birds, the peregrine will sometimes hunt small mammals, small reptiles, or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The peregrine falcon became an endangered species in many areas because of the widespread use of certain pesticides, especially DDT. Since the ban on DDT from the early 1970s, populations have recovered, supported by large-scale protection of nesting places and releases to the wild. The peregrine falcon is a well-respected
falconry Falconry is the hunting of wild animals in their natural state and habitat by means of a trained bird of prey. Small animals are hunted; squirrels and rabbits often fall prey to these birds. Two traditional terms are used to describe a person ...
bird due to its strong hunting ability, high trainability, versatility, and availability via
captive breeding Captive breeding, also known as captive propagation, is the process of plants or animals in controlled environments, such as wildlife reserves, zoos, botanic gardens, and other conservation facilities. It is sometimes employed to help species tha ...
. It is effective on most game bird species, from small to large. It has also been used as a religious, royal, or national symbol across multiple eras and areas of human civilization.


Description

The peregrine falcon has a body length of and a wingspan from . The male and female have similar markings and plumage but, as with many birds of prey, the peregrine falcon displays marked
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
in size, with the female measuring up to 30% larger than the male. Males weigh and the noticeably larger females weigh . In most subspecies, males weigh less than and females weigh more than , and cases of females weighing about 50% more than their male breeding mates are not uncommon. The standard linear measurements of peregrines are: the wing chord measures , the tail measures and the tarsus measures . The back and the long pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies"
below Below may refer to: *Earth * Ground (disambiguation) *Soil *Floor * Bottom (disambiguation) *Less than *Temperatures below freezing *Hell or underworld People with the surname *Ernst von Below (1863–1955), German World War I general *Fred Below ...
); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, coloured like the back but with thin clean bars, is long, narrow, and rounded at the end with a black tip and a white band at the very end. The top of the head and a "moustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the
beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for fo ...
and claws are black. The upper beak is notched near the tip, an
adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
which enables falcons to kill prey by severing the spinal column at the neck. An immature bird is much browner, with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring. A study shows that their black
malar stripe The cheeks ( la, buccae) constitute the area of the face below the eyes and between the nose and the left or right ear. "Buccal" means relating to the cheek. In humans, the region is innervated by the buccal nerve. The area between the inside ...
exists to reduce
glare Glare (derived from GLAss REinforced laminate ) is a fiber metal laminate (FML) composed of several very thin layers of metal (usually aluminum) interspersed with layers of S-2 glass-fiber ''pre-preg'', bonded together with a matrix such as epo ...
from
solar radiation Solar irradiance is the power per unit area ( surface power density) received from the Sun in the form of electromagnetic radiation in the wavelength range of the measuring instrument. Solar irradiance is measured in watts per square metre ...
, allowing them to see better. Photos from The Macaulay Library and iNaturalist showed that the malar stripe is thicker where there is more solar radiation. That supports the solar glare hypothesis.


Taxonomy and systematics

''Falco peregrinus'' was first described under its current
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
by English ornithologist
Marmaduke Tunstall Marmaduke Tunstall (1743 – 11 October 1790) was an English ornithologist and collector. He was the author of ''Ornithologica Britannica'' (1771), probably the first British work to use binomial nomenclature. Tunstall was born at Burton Consta ...
in his 1771 work ''Ornithologia Britannica''. The scientific name ''Falco peregrinus'' is a
Medieval Latin Medieval Latin was the form of Literary Latin used in Roman Catholic Western Europe during the Middle Ages. In this region it served as the primary written language, though local languages were also written to varying degrees. Latin functioned ...
phrase that was used by Albertus Magnus in 1225. The specific name is taken from the fact that juvenile birds were taken while journeying to their breeding location rather than from the nest, as falcon nests were difficult to get at. The Latin term for falcon, , is related to , meaning " sickle", in reference to the silhouette of the falcon's long, pointed wings in flight. The peregrine falcon belongs to a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
whose lineage includes the hierofalcons and the
prairie falcon The prairie falcon (''Falco mexicanus'') is a medium-large sized falcon of western North America. It is about the size of a peregrine falcon or a crow, with an average length of 40 cm (16 in), wingspan of approximately 1 meter (40&n ...
(''F. mexicanus''). This lineage probably diverged from other falcons towards the end of the
Late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene Epoch made up of two stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million years ago) to 5.333 Ma. The ...
or in the
Early Pliocene Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * ...
, about 5–8 
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago ...
(mya). As the peregrine-hierofalcon group includes both
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by thei ...
and North American species, it is likely that the lineage originated in western
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...
or Africa. Its relationship to other falcons is not clear, as the issue is complicated by widespread
hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
confounding mtDNA
sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is called ...
analyses. For example, a genetic lineage of the saker falcon (''F. cherrug'') is known which originated from a male saker producing fertile young with a female peregrine ancestor, and the descendants further breeding with sakers. Today, peregrines are regularly paired in captivity with other species such as the lanner falcon (''F. biarmicus'') to produce the "
perilanner The word perilanner is a falconer's term for a hybrid between a peregrine falcon and a lanner falcon. It is larger and faster than a lanner, but does not fly as far as a peregrine and thus is less likely to fly far away and become lost. As a rule ...
", a somewhat popular bird in
falconry Falconry is the hunting of wild animals in their natural state and habitat by means of a trained bird of prey. Small animals are hunted; squirrels and rabbits often fall prey to these birds. Two traditional terms are used to describe a person ...
as it combines the peregrine's hunting skill with the lanner's hardiness, or the gyrfalcon to produce large, strikingly coloured birds for the use of falconers. As can be seen, the peregrine is still genetically close to the hierofalcons, though their lineages diverged in the
Late Pliocene Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effe ...
(maybe some 2.5–2 mya in the Gelasian).


Subspecies

Numerous
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
of ''Falco peregrinus'' have been described, with 19 accepted by the 1994 '' Handbook of the Birds of the World'', which considers the Barbary falcon of the
Canary Islands The Canary Islands (; es, :es:Canarias, Canarias, ), also known informally as the Canaries, are a Spanish Autonomous communities of Spain, autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to ...
and coastal
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
to be two subspecies (''pelegrinoides'' and ''babylonicus'') of ''Falco peregrinus'', rather than a distinct species, ''F. pelegrinoides''. The following map shows the general ranges of these 19 subspecies. *', described by Bonaparte in 1838, is known as the American peregrine falcon or "duck hawk"; its scientific name means "duck peregrine falcon". At one time, it was partly included in ''leucogenys''. It is mainly found in the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico ...
today. It was formerly common throughout North America between the tundra and northern
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
, where current
reintroduction Species reintroduction is the deliberate release of a species into the wild, from captivity or other areas where the organism is capable of survival. The goal of species reintroduction is to establish a healthy, genetically diverse, self-sustainin ...
efforts are seeking to restore the population. Most mature ''anatum'', except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory '' tundrius'', only considered distinct since 1968. It is similar to the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
, but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh , while females weigh . It has become extinct in eastern North America and populations there are hybrids as a result of reintroductions of birds from elsewhere. *''Falco peregrinus babylonicus'', described by P.L. Sclater in 1861, is found in eastern
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
along the Hindu Kush and the
Tian Shan The Tian Shan,, , otk, 𐰴𐰣 𐱅𐰭𐰼𐰃, , tr, Tanrı Dağı, mn, Тэнгэр уул, , ug, تەڭرىتاغ, , , kk, Тәңіртауы / Алатау, , , ky, Теңир-Тоо / Ала-Тоо, , , uz, Tyan-Shan / Tangritog‘ ...
to the
Mongolian Altai The Altai Mountains (), also spelled Altay Mountains, are a mountain range in Central and East Asia, where Russia, China, Mongolia and Kazakhstan converge, and where the rivers Irtysh and Ob have their headwaters. The massif merges with the S ...
ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than ''pelegrinoides'' and somewhat similar to a small, pale lanner falcon (''Falco biarmicus''). Males weigh , while females weigh . *', described by Sharpe in 1873, is also known as the Mediterranean peregrine falcon or the Maltese falcon. It includes ''caucasicus'' and most specimens of the proposed race ''punicus'', though others may be ''pelegrinoides'' ( Barbary falcons), or perhaps the rare hybrids between these two which might occur around
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
. They occur from the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
around the Mediterranean, except in
arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ...
regions, to the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...
. They are non-migratory. It is smaller than the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
and the underside usually has a rusty hue. Males weigh around , while females weigh up to . *', described by John Latham in 1790, it was formerly called ''leucogenys'' and includes ''caeruleiceps''. It breeds in the
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
tundra of Eurasia from
Murmansk Oblast Murmansk Oblast (russian: Му́рманская о́бласть, p=ˈmurmənskəjə ˈobləsʲtʲ, r=Murmanskaya oblast, ''Murmanskaya oblast''; Kildin Sami: Мурман е̄ммьне, ''Murman jemm'ne'') is a federal subject (an oblast) o ...
to roughly Yana and Indigirka Rivers,
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
. It is completely migratory and travels south in winter as far as
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;; ...
and
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
. It is often seen around wetland habitats. It is paler than the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
, especially on the crown. Males weigh , while females weigh . *''Falco peregrinus cassini'', described by Sharpe in 1873, is also known as the austral peregrine falcon. It includes ''kreyenborgi'', the pallid falcon, a leucistic colour morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
through
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
, northern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the List of South American countries by area, second-largest ...
and
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
to
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla ...
and the
Falkland Islands The Falkland Islands (; es, Islas Malvinas, link=no ) is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and about from Cape Dubouze ...
. It is non-migratory. It is similar to the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
, but slightly smaller with a black ear region. The pallid falcon morph ''kreyenborgi'' is medium grey above, has little barring below and has a head pattern like the saker falcon (''Falco cherrug''), but the ear region is white. *''Falco peregrinus ernesti'', described by Sharpe in 1894, is found from the Sunda Islands to the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
and south to eastern
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torres ...
and the nearby
Bismarck Archipelago The Bismarck Archipelago (, ) is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea. Its area is about 50,000 square km. History The first inhabitants o ...
. Its geographical separation from '' nesiotes'' requires confirmation. It is non-migratory. It differs from the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
in the very dark, dense barring on its underside and its black ear coverts. *''Falco peregrinus furuitii'', described by Momiyama in 1927, is found on the Izu and
Ogasawara Islands The Bonin Islands, also known as the , are an archipelago of over 30 subtropical and tropical islands, some directly south of Tokyo, Japan and northwest of Guam. The name "Bonin Islands" comes from the Japanese word ''bunin'' (an archaic read ...
south of
Honshū , historically called , is the largest and most populous island of Japan. It is located south of Hokkaidō across the Tsugaru Strait, north of Shikoku across the Inland Sea, and northeast of Kyūshū across the Kanmon Straits. The island sepa ...
, Japan. It is non-migratory. It is very rare and may only remain on a single island. It is a dark form, resembling '' pealei'' in colour, but darker, especially on the tail. *''Falco peregrinus japonensis'', described by Gmelin in 1788, includes ''kleinschmidti'', ''pleskei'', and ''harterti'', and seems to refer to intergrades with '' calidus''. It is found from northeast
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
to Kamchatka (though it is possibly replaced by ''pealei'' on the coast there) and
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...
. Northern populations are migratory, while those of Japan are resident. It is similar to the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
, but the young are even darker than those of '' anatum''. *''Falco peregrinus macropus'', described by Swainson in 1837, is the Australian peregrine falcon. It is found in
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
in all regions except the southwest. It is non-migratory. It is similar to '' brookei'' in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large. *''Falco peregrinus madens'', described by
Ripley Ripley may refer to: People and characters * Ripley (name) * ''Ripley'', the test mannequin aboard the first International Space Station space station Dragon 2 space test flight Crew Dragon Demo-1 * Ellen Ripley, a fictional character from the Al ...
and Watson in 1963, is unusual in having some
sexual dichromatism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
. If the Barbary falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands and is non-migratory; it is also endangered, with only six to eight pairs surviving. Males have a rufous wash on the crown, nape, ears and back; the underside is conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape. *''Falco peregrinus minor'', first described by Bonaparte in 1850. It was formerly often known as ''perconfusus''. It is sparsely and patchily distributed throughout much of
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
and widespread in
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number o ...
. It apparently reaches north along the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
coast as far as
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to A ...
. It is non-migratory and dark-coloured. This is the smallest subspecies, with smaller males weighing as little as approximately . *', described by Mayr in 1941, is found in
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consis ...
and probably also
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of ...
and
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
. It is non-migratory. *', described by Ridgway in 1873, is
Peale's falcon Peale's falcon (''Falco peregrinus pealei'') is a subspecies of the peregrine falcon. This race was first identified by the ornithologist Robert Ridgway in 1873, named in honor of Titian Ramsay Peale. These birds are the largest subspecies of pe ...
and includes ''rudolfi''. It is found in the
Pacific Northwest The Pacific Northwest (sometimes Cascadia, or simply abbreviated as PNW) is a geographic region in western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Thou ...
of North America, northwards from
Puget Sound Puget Sound ( ) is a sound of the Pacific Northwest, an inlet of the Pacific Ocean, and part of the Salish Sea. It is located along the northwestern coast of the U.S. state of Washington. It is a complex estuarine system of interconnected m ...
along the
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, for ...
coast (including the Haida Gwaii), along the Gulf of Alaska and the
Aleutian Islands The Aleutian Islands (; ; ale, Unangam Tanangin,”Land of the Aleuts", possibly from Chukchi ''aliat'', "island"), also called the Aleut Islands or Aleutic Islands and known before 1867 as the Catherine Archipelago, are a chain of 14 large v ...
to the far eastern
Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Am ...
coast of
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
, and may also occur on the
Kuril Islands The Kuril Islands or Kurile Islands (; rus, Кури́льские острова́, r=Kuril'skiye ostrova, p=kʊˈrʲilʲskʲɪjə ɐstrɐˈva; Japanese language, Japanese: or ) are a volcanic archipelago currently administered as part of Sakh ...
and the coasts of Kamchatka. It is non-migratory. It is the largest subspecies and it looks like an oversized and darker '' tundrius'' or like a strongly barred and large '' anatum''. The bill is very wide. Juveniles occasionally have pale crowns. Males weigh , while females weigh . *'' Falco peregrinus pelegrinoides'', first described by
Temminck Coenraad Jacob Temminck (; 31 March 1778 – 30 January 1858) was a Dutch aristocrat, zoologist and museum director. Biography Coenraad Jacob Temminck was born on 31 March 1778 in Amsterdam in the Dutch Republic. From his father, Jacob Temminc ...
in 1829, is found in the
Canary Islands The Canary Islands (; es, :es:Canarias, Canarias, ), also known informally as the Canaries, are a Spanish Autonomous communities of Spain, autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to ...
through North Africa and the
Near East The ''Near East''; he, המזרח הקרוב; arc, ܕܢܚܐ ܩܪܒ; fa, خاور نزدیک, Xāvar-e nazdik; tr, Yakın Doğu is a geographical term which roughly encompasses a transcontinental region in Western Asia, that was once the hist ...
to
Mesopotamia Mesopotamia ''Mesopotamíā''; ar, بِلَاد ٱلرَّافِدَيْن or ; syc, ܐܪܡ ܢܗܪ̈ܝܢ, or , ) is a historical region of Western Asia situated within the Tigris–Euphrates river system, in the northern part of the ...
. It is most similar to '' brookei'', but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
; females weigh around . *'' Falco peregrinus peregrinator'', described by
Sundevall Carl Jakob Sundevall (22 October 1801, Högestad – 2 February 1875) was a Swedish zoologist. Sundevall studied at Lund University, where he became a Ph.D. in 1823. After traveling to East Asia, he studied medicine, graduating as Doctor of Me ...
in 1837, is known as the Indian peregrine falcon, black shaheen, Indian shaheen or shaheen falcon. It was formerly sometimes known as ''Falco atriceps'' or ''Falco shaheen''. Its range includes
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;; ...
from across the
Indian subcontinent The Indian subcontinent is a physiographical region in Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian Ocean from the Himalayas. Geopolitically, it includes the countries of Bangladesh, Bhutan, In ...
to
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
and southeastern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
. In India, the shaheen falcon is reported from all states except
Uttar Pradesh Uttar Pradesh (; , 'Northern Province') is a state in northern India. With over 200 million inhabitants, it is the most populated state in India as well as the most populous country subdivision in the world. It was established in 195 ...
, mainly from rocky and hilly regions. The shaheen falcon is also reported from the
Andaman and Nicobar Islands The Andaman and Nicobar Islands is a union territory of India consisting of 572 islands, of which 37 are inhabited, at the junction of the Bay of Bengal and the Andaman Sea. The territory is about north of Aceh in Indonesia and separated f ...
in the
Bay of Bengal The Bay of Bengal is the northeastern part of the Indian Ocean, bounded on the west and northwest by India, on the north by Bangladesh, and on the east by Myanmar and the Andaman and Nicobar Islands of India. Its southern limit is a line bet ...
. It has a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on man-made structures such as buildings and cellphone transmission towers. A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory and is small and dark, with rufous underparts. In
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
this species is found to favour the higher hills, while the migrant '' calidus'' is more often seen along the coast. *', the nominate (first-named) subspecies, described by Tunstall in 1771, breeds over much of temperate
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...
between the
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless mou ...
in the north and the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
,
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...
region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh , while females weigh . It includes ''brevirostris'', ''germanicus'', ''rhenanus'' and ''riphaeus''. *''Falco peregrinus radama'', described by
Hartlaub Karel Johan Gustav Hartlaub (8 November 1814 – 29 November 1900) was a German physician and ornithologist. Hartlaub was born in Bremen, and studied at Bonn and Berlin before graduating in medicine at Göttingen. In 1840, he began to study and c ...
in 1861, is found in
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Afric ...
and the
Comoros The Comoros,, ' officially the Union of the Comoros,; ar, الاتحاد القمري ' is an independent country made up of three islands in southeastern Africa, located at the northern end of the Mozambique Channel in the Indian Ocean. It ...
. It is non-migratory. *''Falco peregrinus submelanogenys'', described by Mathews in 1912, is the Southwest Australian peregrine falcon. It is found in southwestern Australia and is non-migratory. *', described by C.M. White in 1968, was at one time included in ''leucogenys''. It is found in the Arctic tundra of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
to
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
, and migrates to wintering grounds in
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known a ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
. Most vagrants that reach western Europe belong to this subspecies, which was previously considered synonymous with '' anatum''. It is the New World equivalent to '' calidus''. It is smaller and paler than ''anatum''; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in ''calidus''. Juveniles are browner and less grey than in ''calidus'' and paler, sometimes almost sandy, than in ''anatum''. Males weigh , while females weigh . Despite its current recognition as a valid subspecies, a population genetic study of both pre-decline (i.e., museum) and recovered contemporary populations failed to distinguish genetically the ''anatum'' and ''tundrius'' subspecies.


Barbary falcon

The Barbary falcon is a subspecies of the peregrine falcon that inhabits parts of North Africa; namely, from the Canary Islands to the Arabian Peninsula. There is discussion concerning the taxonomic status of the bird, with some considering it a subspecies of the peregrine falcon and others considering it a full species with two subspecies (White et al. 2013). Compared to the other peregrine falcon subspecies, Barbary falcons sport a slimmer body and a distinct plumage color pattern. Despite numbers and range of these birds throughout the Canary Islands generally increasing, they are considered endangered, with human interference through falconry and shooting threatening their well-being. Falconry can further complicate the speciation and genetics of these Canary Islands falcons, as the practice promotes genetic mixing between individuals from outside the islands with those originating from the islands. Population density of the Barbary falcons on Tenerife, the biggest of the seven major Canary Islands, was found to be 1.27 pairs/100 km², with the mean distance between pairs being 5869 ± 3338 m. The falcons were only observed near large and natural cliffs with a mean altitude of 697.6 m. Falcons show an affinity for tall cliffs away from human-mediated establishments and presence. Barbary falcons have a red neck patch, but otherwise differ in appearance from the peregrine falcon proper merely according to Gloger's rule, relating pigmentation to environmental humidity. The Barbary falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the peregrine falcon, but less often and far less pronounced. The Barbary falcon's shoulder and
pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...
bones are stout by comparison with the peregrine falcon and its feet are smaller. Barbary falcons breed at different times of year than neighboring peregrine falcon subspecies, but they are capable of interbreeding. There is a 0.6–0.7% genetic distance in the peregrine falcon-Barbary falcon ("peregrinoid") complex.


Ecology and behaviour

The peregrine falcon lives mostly along
mountain range A mountain range or hill range is a series of mountains or hills arranged in a line and connected by high ground. A mountain system or mountain belt is a group of mountain ranges with similarity in form, structure, and alignment that have ari ...
s, river valleys,
coastline The coast, also known as the coastline or seashore, is defined as the area where land meets the ocean, or as a line that forms the boundary between the land and the coastline. The Earth has around of coastline. Coasts are important zones in ...
s, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologi ...
s typically migrate great distances during the northern winter. The peregrine falcon reaches faster speeds than any other animal on the planet when performing the stoop, which involves soaring to a great height and then diving steeply at speeds of over , hitting one wing of its prey so as not to harm itself on impact. The air pressure from such a dive could possibly damage a bird's
lung The lungs are the primary organs of the respiratory system in humans and most other animals, including some snails and a small number of fish. In mammals and most other vertebrates, two lungs are located near the backbone on either side of ...
s, but small bony tubercles on a falcon's nostrils are theorized to guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their
nictitating membrane The nictitating membrane (from Latin '' nictare'', to blink) is a transparent or translucent third eyelid present in some animals that can be drawn across the eye from the medial canthus to protect and moisten it while maintaining vision. All ...
s (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. The distinctive malar stripe or 'moustache', a dark area of feathers below the eyes, is thought to reduce solar glare and improve contrast sensitivity when targeting fast moving prey in bright light condition; the malar stripe has been found to be wider and more pronounced in regions of the world with greater solar radiation supporting this solar glare hypothesis. Peregrine falcons have a flicker fusion frequency of 129 Hz (cycles per second), very fast for a bird of its size, and much faster than mammals. A study testing the flight physics of an "ideal falcon" found a theoretical speed limit at for low-altitude flight and for high-altitude flight. In 2005, Ken Franklin recorded a falcon stooping at a top speed of . The life span of peregrine falcons in the wild is up to 19 years 9 months. Mortality in the first year is 59–70%, declining to 25–32% annually in adults. Apart from such
anthropogenic Anthropogenic ("human" + "generating") is an adjective that may refer to: * Anthropogeny, the study of the origins of humanity Counterintuitively, anthropogenic may also refer to things that have been generated by humans, as follows: * Human i ...
threats as collision with human-made objects, the peregrine may be killed by larger
hawk Hawks are birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. This subfa ...
s and owls. The peregrine falcon is host to a range of parasites and
pathogen In biology, a pathogen ( el, πάθος, "suffering", "passion" and , "producer of") in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a g ...
s. It is a vector for Avipoxvirus,
Newcastle disease virus Newcastle usually refers to: *Newcastle upon Tyne, a city and metropolitan borough in Tyne and Wear, England *Newcastle-under-Lyme, a town in Staffordshire, England *Newcastle, New South Wales, a metropolitan area in Australia, named after Newcastle ...
, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some
mycoses Fungal infection, also known as mycosis, is disease caused by fungi. Different types are traditionally divided according to the part of the body affected; superficial, subcutaneous, and systemic. Superficial fungal infections include common ...
and bacterial infections. Endoparasites include '' Plasmodium relictum'' (usually not causing
malaria Malaria is a mosquito-borne infectious disease that affects humans and other animals. Malaria causes symptoms that typically include fever, tiredness, vomiting, and headaches. In severe cases, it can cause jaundice, seizures, coma, or death. ...
in the peregrine falcon), Strigeidae
trematode Trematoda is a class of flatworms known as flukes. They are obligate internal parasites with a complex life cycle requiring at least two hosts. The intermediate host, in which asexual reproduction occurs, is usually a snail. The definitive h ...
s, '' Serratospiculum amaculata'' (
nematode The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant- parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a bro ...
), and
tapeworm Eucestoda, commonly referred to as tapeworms, is the larger of the two subclasses of flatworms in the class Cestoda (the other subclass is Cestodaria). Larvae have six posterior hooks on the scolex (head), in contrast to the ten-hooked Cesto ...
s. Known peregrine falcon
ectoparasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson ha ...
s are
chewing lice The Mallophaga are a possibly paraphyletic section of lice Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been reco ...
, ''
Ceratophyllus garei ''Ceratophyllus garei'', the duck flea, is a species of flea in the family Ceratophyllidae. It was described by Rothschild Rothschild () is a name derived from the German ''zum rothen Schild'' (with the old spelling "th"), meaning "with the r ...
'' (a
flea Flea, the common name for the order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by ingesting the blood of their hosts. Adult fleas grow to about long, ...
), and Hippoboscidae flies (''
Icosta nigra ''Icosta'' are genus of biting flies in the family of louse flies, Hippoboscidae. There are 52 known species, making it the largest Hippoboscid genus. All species are parasites of birds. Distribution Worldwide, excluding Antarctica. Systematic ...
'', ''
Ornithoctona erythrocephala ''Ornithoctona'' are genus of biting flies in the family of louse flies, Hippoboscidae. There are 12 known species. All species are parasites of birds. Distribution ''Ornithoctona'' are found worldwide with the exception of Antarctica. Systemat ...
''). In the Arctic Peregrine falcons chasing away small rodent predators from their nesting territory and Rough-legged Hawks (''Buteo lagopus'') could use these hot spots as a nesting territory.


Feeding

The peregrine falcon feeds almost exclusively on medium-sized birds such as pigeons and doves,
waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which ...
, songbirds, and
wader 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
s. This falcon tends to nest on tall buildings or bridges, and these urban dwelling birds subsist mostly on different pigeons. Worldwide, it is estimated that between 1,500 and 2,000 bird species (up to roughly a fifth of the world's bird species) are predated somewhere by these falcons. In North America, prey has varied in size from
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With about 361 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are ...
s ('' Selasphorus'' and ''
Archilochus Archilochus (; grc-gre, Ἀρχίλοχος ''Arkhilokhos''; c. 680 – c. 645 BC) was a Greek lyric poet of the Archaic period from the island of Paros. He is celebrated for his versatile and innovative use of poetic meters, and is the ...
'' ssp.) to a sandhill crane (killed in Alaska by a peregrine in a stoop), although most prey taken by peregrines weigh from (small
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
s) to (such as
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form ...
s and
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century ...
s). The peregrine falcon takes the most diverse range of bird species of any raptor in North America, with more than 300 species having fallen victim to the falcon, including nearly 100
shorebird 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
s. Smaller
hawk Hawks are birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. This subfa ...
s and owls are regularly predated, mainly smaller falcons such as the American kestrel,
merlin Merlin ( cy, Myrddin, kw, Marzhin, br, Merzhin) is a mythical figure prominently featured in the legend of King Arthur and best known as a mage, with several other main roles. His usual depiction, based on an amalgamation of historic and leg ...
and sharp-shinned hawks. In urban areas, the main component of the peregrine's diet is the rock or
feral pigeon Feral pigeons (''Columba livia domestica'' or ''Columba livia forma urbana''), also called city doves, city pigeons, or street pigeons,Nagy, Kelsi, and Johnson, Phillip David. ''Trash animals: how we live with natures filthy, feral, invasive, an ...
, which comprise 80% or more of the dietary intake for peregrines in some cities. Other common city birds are also taken regularly, including mourning doves, common wood pigeons, common swifts,
northern flicker The northern flicker or common flicker (''Colaptes auratus'') is a medium-sized bird of the woodpecker family. It is native to most of North America, parts of Central America, Cuba, and the Cayman Islands, and is one of the few woodpecker spec ...
s,
common starling The common starling or European starling (''Sturnus vulgaris''), also known simply as the starling in Great Britain and Ireland, is a medium-sized passerine bird in the starling family, Sturnidae. It is about long and has glossy black plumage ...
s,
American robin The American robin (''Turdus migratorius'') is a migratory bird of the true thrush genus and Turdidae, the wider thrush family. It is named after the European robin because of its reddish-orange breast, though the two species are not closely ...
s,
common blackbird The common blackbird (''Turdus merula'') is a species of true thrush. It is also called the Eurasian blackbird (especially in North America, to distinguish it from the unrelated New World blackbirds), or simply the blackbird where this does not ...
s, and corvids (such as magpies or
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
,
house A house is a single-unit residential building. It may range in complexity from a rudimentary hut to a complex structure of wood, masonry, concrete or other material, outfitted with plumbing, electrical, and heating, ventilation, and air cond ...
, and American crows). Other than bats taken at night, the peregrine rarely hunts mammals, but will on occasion take small species such as rats,
vole Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...
s,
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The g ...
s,
shrew Shrews (family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to differ ...
s,
mice A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus'' ...
and
squirrel Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrels. ...
s. Coastal populations of the large subspecies ''pealei'' feed almost exclusively on
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s. In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies ''tundrius'' was observed while successfully hunting a juvenile
scarlet ibis The scarlet ibis (''Eudocimus ruber'') is a species of ibis in the bird family Threskiornithidae. It inhabits tropical South America and part of the Caribbean. In form, it resembles most of the other twenty-seven extant species of ibis, but ...
. Insects and reptiles make up a small proportion of the diet, which varies greatly depending on what prey is available. The peregrine falcon hunts most often at dawn and dusk, when prey are most active, but also nocturnally in cities, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by peregrines include species as diverse as yellow-billed cuckoo, black-necked grebe, virginia rail, and common quail. The peregrine requires open space in order to hunt, and therefore often hunts over open water,
marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found ...
es,
valley A valley is an elongated low area often running between hills or mountains, which will typically contain a river or stream running from one end to the other. Most valleys are formed by erosion of the land surface by rivers or streams ove ...
s, fields, and
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless mou ...
, searching for prey either from a high perch or from the air. Large congregations of migrants, especially species that gather in the open like shorebirds, can be quite attractive to hunting peregrines. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. Prey is typically struck and captured in mid-air; the peregrine falcon strikes its prey with a clenched foot, stunning or killing it with the impact, then turns to catch it in mid-air. If its prey is too heavy to carry, a peregrine will drop it to the ground and eat it there. If they miss the initial strike, peregrines will chase their prey in a twisting flight. Although previously thought rare, several cases of peregrines contour-hunting, i.e. using natural contours to surprise and ambush prey on the ground, have been reported and even rare cases of prey being pursued on foot. In addition, peregrines have been documented preying on chicks in nests, from birds such as kittiwakes. Prey is plucked before consumption. A recent study showed the presence of peregrines benefits non-preferred species while at the same time causing a decline in its preferred prey. As of 2018, the fastest recorded falcon was at 242 mph (nearly 390 km/h). Researchers at the University of Groningen in the Netherlands and at Oxford University used 3D computer simulations in 2018 to show that the high speed allows peregrines to gain better maneuverability and precision in strikes."Falcon Attack: How Peregrine Falcons Maneuver At Nearly 225 MPH"
Fiona McMillan, 13 April 2018, ''Forbes''


Reproduction

The peregrine falcon is sexually mature at one to three years of age, but in larger populations they breed after two to three years of age. A pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons. During the breeding season, the peregrine falcon is territorial; nesting pairs are usually more than apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two up to seven in a 16-year period. The peregrine falcon nests in a scrape, normally on cliff edges. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. Cliff nests are generally located under an overhang, on ledges with vegetation. South-facing sites are favoured. In some regions, as in parts of
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
and on the west coast of northern North America, large tree hollows are used for nesting. Before the demise of most European peregrines, a large population of peregrines in central and western Europe used the disused nests of other large birds. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. In many parts of its range, peregrines now also nest regularly on tall buildings or bridges; these human-made structures used for breeding closely resemble the natural cliff ledges that the peregrine prefers for its nesting locations. The pair defends the chosen nest site against other peregrines, and often against ravens,
heron The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychu ...
s, and
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century ...
s, and if ground-nesting, also such mammals as foxes,
wolverine The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for " glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is a musc ...
s, felids,
bear Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the No ...
s,
wolves The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly un ...
, and mountain lions. Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, j ...
s, large owls, or gyrfalcons. The most serious predators of peregrine nests in North America and Europe are the great horned owl and the
Eurasian eagle owl The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl that resides in much of Eurasia. It is also called the Uhu and it is occasionally abbreviated to just the eagle-owl in Europe. It is one of the largest species of owl, and female ...
. When reintroductions have been attempted for peregrines, the most serious impediments were these two species of owls routinely picking off nestlings, fledglings and adults by night. Peregrines defending their nests have managed to kill raptors as large as
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird ...
s and
bald eagle The bald eagle (''Haliaeetus leucocephalus'') is a bird of prey found in North America. A sea eagle, it has two known subspecies and forms a species pair with the white-tailed eagle (''Haliaeetus albicilla''), which occupies the same niche as ...
s (both of which they normally avoid as potential predators) that have come too close to the nest by ambushing them in a full stoop. In one instance, when a snowy owl killed a newly fledged peregrine, the larger owl was in turn killed by a stooping peregrine parent. The date of egg-laying varies according to locality, but is generally from February to March in the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
, and from July to August in the Southern Hemisphere, although the Australian subspecies ''macropus'' may breed as late as November, and
equator The equator is a circle of latitude, about in circumference, that divides Earth into the Northern and Southern hemispheres. It is an imaginary line located at 0 degrees latitude, halfway between the North and South poles. The term can also ...
ial populations may nest anytime between June and December. If the eggs are lost early in the nesting season, the female usually lays another clutch, although this is extremely rare in the Arctic due to the short summer season. Generally three to four eggs, but sometimes as few as one or as many as five, are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female, with the male also helping with the incubation of the eggs during the day, but only the female incubating them at night. The average number of young found in nests is 2.5, and the average number that fledge is about 1.5, due to the occasional production of infertile eggs and various natural losses of nestlings. After hatching, the chicks (called "es") are covered with creamy-white down and have disproportionately large feet. The male (called the "") and the female (simply called the "falcon") both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.


Relationship with humans


Use in falconry

The peregrine falcon is a highly admired falconry bird, and has been used in
falconry Falconry is the hunting of wild animals in their natural state and habitat by means of a trained bird of prey. Small animals are hunted; squirrels and rabbits often fall prey to these birds. Two traditional terms are used to describe a person ...
for more than 3,000 years, beginning with nomads in
central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the fo ...
. Its advantages in falconry include not only its athleticism and eagerness to hunt, but an equable disposition that leads to it being one of the easier falcons to train. The peregrine falcon has the additional advantage of a natural flight style of circling above the falconer ("waiting on") for game to be flushed, and then performing an effective and exciting high-speed diving stoop to take the quarry. The speed of the stoop not only allows the falcon to catch fast flying birds, it also enhances the falcon's ability to exert maneuvers to catch highly agile prey, and allows the falcon to deliver a knockout blow with a fist-like clenched talon against game that may be much larger than itself. Additionally the versatility of the species, with agility allowing capture of smaller birds and a strength and attacking style allowing capture of game much larger than themselves, combined with the wide size range of the many peregrine subspecies, means there is a subspecies suitable to almost any size and type of game bird. This size range, evolved to fit various environments and prey species, is from the larger females of the largest subspecies to the smaller males of the smallest subspecies, approximately five to one (approximately 1500 g to 300 g). The males of smaller and medium-sized subspecies, and the females of the smaller subspecies, excel in the taking of swift and agile small game birds such as dove, quail, and smaller ducks. The females of the larger subspecies are capable of taking large and powerful game birds such as the largest of duck species, pheasant, and grouse. Peregrine falcons handled by falconers are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety. They were also used to intercept homing pigeons during World War II. Peregrine falcons have been successfully bred in captivity, both for falconry and for release into the wild. Until 2004 nearly all peregrines used for falconry in the US were captive-bred from the progeny of falcons taken before the US Endangered Species Act was enacted and from those few infusions of wild genes available from Canada and special circumstances. Peregrine falcons were removed from the United States' endangered species list in 1999. The successful recovery program was aided by the effort and knowledge of falconers – in collaboration with
The Peregrine Fund The Peregrine Fund (named after the bird of prey of the same name the peregrine falcon) is a non-profit organization founded in 1970 that conserves threatened and endangered birds of prey worldwide. The successful recovery of the peregrine falcon ...
and state and federal agencies – through a technique called hacking. Finally, after years of close work with the US Fish and Wildlife Service, a limited take of wild peregrines was allowed in 2004, the first wild peregrines taken specifically for falconry in over 30 years. The development of captive breeding methods has led to peregrines being commercially available for falconry use, thus mostly eliminating the need to capture wild birds for support of falconry. The main reason for taking wild peregrines at this point is to maintain healthy genetic diversity in the breeding lines. Hybrids of peregrines and gyrfalcons are also available that can combine the best features of both species to create what many consider to be the ultimate falconry bird for the taking of larger game such as the sage-grouse. These hybrids combine the greater size, strength, and horizontal speed of the gyrfalcon with the natural propensity to stoop and greater warm weather tolerance of the peregrine.


Decline due to pesticides

The peregrine falcon became an endangered species over much of its range because of the use of
organochlorine pesticide An organochloride, organochlorine compound, chlorocarbon, or chlorinated hydrocarbon is an organic compound containing at least one covalently bonded atom of chlorine. The chloroalkane class (alkanes with one or more hydrogens substituted by chlor ...
s, especially DDT, during the 1950s, '60s, and '70s. Pesticide biomagnification caused organochlorine to build up in the falcons' fat tissues, reducing the amount of calcium in their eggshells. With thinner shells, fewer falcon eggs survived until hatching. In addition, the PCB concentrations found in these falcons is dependent upon the age of the falcon. While high levels are still found in young birds (only a few months old) and even higher concentrations are found in more mature falcons, further increasing in adult peregrine falcons. These pesticides caused falcon prey to also have thinner eggshells (one example of prey being the Black Petrels). In several parts of the world, such as the eastern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
and
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to ...
, this species became extirpated (locally extinct) as a result. An alternate point of view is that populations in the eastern North America had vanished due to hunting and egg collection. Following the ban of organochlorine pesticides, the reproductive success of Peregrines increased in Scotland in terms of territory occupancy and breeding success, although spatial variation in recovery rates indicate that in some areas Peregrines were also impacted by other factors such as persecution.


Recovery efforts

Peregrine falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a peregrine's head, so they cannot see to
imprint Imprint or imprinting may refer to: Entertainment * ''Imprint'' (TV series), Canadian television series * "Imprint" (''Masters of Horror''), episode of TV show ''Masters of Horror'' * ''Imprint'' (film), a 2007 independent drama/thriller film ...
on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced, forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment. Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The peregrine falcon was removed from the U.S.
Endangered Species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
list on 25 August 1999. Some controversy has existed over the origins of captive breeding stock used by
the Peregrine Fund The Peregrine Fund (named after the bird of prey of the same name the peregrine falcon) is a non-profit organization founded in 1970 that conserves threatened and endangered birds of prey worldwide. The successful recovery of the peregrine falcon ...
in the recovery of peregrine falcons throughout the contiguous United States. Several peregrine subspecies were included in the breeding stock, including birds of Eurasian origin. Due to the extirpation of the eastern population of ''Falco peregrinus anatum'', the near-extirpation of ''anatum'' in the Midwest and the limited gene pool within North American breeding stock, the inclusion of non-native
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
was justified to optimize the
genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species, it ranges widely from the number of species to differences within species and can be attributed to the span of survival for a species. It is dis ...
found within the species as a whole. During the 1970s, peregrine falcons in Finland experienced a
population bottleneck A population bottleneck or genetic bottleneck is a sharp reduction in the size of a population due to environmental events such as famines, earthquakes, floods, fires, disease, and droughts; or human activities such as specicide, widespread violen ...
as a result of large declines associated with
bio-accumulation Bioaccumulation is the gradual accumulation of substances, such as pesticides or other chemicals, in an organism. Bioaccumulation occurs when an organism absorbs a substance at a rate faster than that at which the substance is lost or eliminated ...
of organochloride pesticides. However, the genetic diversity of peregrines in Finland is similar to other populations, indicating that high dispersal rates have maintained the genetic diversity of this species. Since peregrine falcon eggs and chicks are still often targeted by illegal poachers, it is common practice not to publicize unprotected nest locations.


Current status

Populations of the peregrine falcon have bounced back in most parts of the world. In the United Kingdom, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the
Royal Society for the Protection of Birds The Royal Society for the Protection of Birds (RSPB) is a charitable organisation registered in England and Wales and in Scotland. It was founded in 1889. It works to promote conservation and protection of birds and the wider environment thro ...
. The RSPB has estimated that there are 1,402 breeding pairs in the UK. In Canada, where peregrines were identified as endangered in 1978 (in the
Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
territory of northern
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...
that year, only a single breeding pair was identified), the
Committee on the Status of Endangered Wildlife in Canada The Committee on the Status of Endangered Wildlife in Canada (COSEWIC, French: Comité sur la situation des espèces en péril au Canada, COSEPAC) is an independent committee of wildlife experts and scientists whose "raison d'être is to identify s ...
declared the species no longer at risk in December 2017. Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalising on the urban
feral pigeon Feral pigeons (''Columba livia domestica'' or ''Columba livia forma urbana''), also called city doves, city pigeons, or street pigeons,Nagy, Kelsi, and Johnson, Phillip David. ''Trash animals: how we live with natures filthy, feral, invasive, an ...
populations for food. In
Southampton Southampton () is a port city in the ceremonial county of Hampshire in southern England. It is located approximately south-west of London and west of Portsmouth. The city forms part of the South Hampshire built-up area, which also covers Po ...
, a nest prevented restoration of mobile telephony services for several months, after
Vodafone Vodafone Group plc () is a British multinational telecommunications company. Its registered office and global headquarters are in Newbury, Berkshire, England. It predominantly operates services in Asia, Africa, Europe, and Oceania. , Vod ...
engineers despatched to repair a faulty
transmitter mast Radio masts and towers are typically tall structures designed to support antennas for telecommunications and broadcasting, including television. There are two main types: guyed and self-supporting structures. They are among the tallest human-mad ...
discovered a nest in the mast, and were prevented by the Wildlife and Countryside Act – on pain of a possible prison sentence – from proceeding with repairs until the chicks fledged. In many parts of the world peregrine falcons have adapted to urban habitats, nesting on
cathedral A cathedral is a church that contains the ''cathedra'' () of a bishop, thus serving as the central church of a diocese, conference, or episcopate. Churches with the function of "cathedral" are usually specific to those Christian denominations ...
s,
skyscraper A skyscraper is a tall continuously habitable building having multiple floors. Modern sources currently define skyscrapers as being at least or in height, though there is no universally accepted definition. Skyscrapers are very tall high-ri ...
window ledges, tower blocks, and the towers of
suspension bridge A suspension bridge is a type of bridge in which the deck is hung below suspension cables on vertical suspenders. The first modern examples of this type of bridge were built in the early 1800s. Simple suspension bridges, which lack vertical ...
s. Many of these nesting birds are encouraged, sometimes gathering media attention and often monitored by cameras.


Cultural significance

Due to its striking hunting technique, the peregrine has often been associated with aggression and martial prowess. The Ancient Egyptian solar deity Ra was often represented as a man with the head of a Peregrine Falcon adorned with the solar disk. Native Americans of the Mississippian culture (c. 800–1500) used the peregrine, along with several other birds of prey, in imagery as a symbol of "aerial (celestial) power" and buried men of high status in costumes associating to the ferocity of raptorial birds. In the
late Middle Ages The Late Middle Ages or Late Medieval Period was the period of European history lasting from AD 1300 to 1500. The Late Middle Ages followed the High Middle Ages and preceded the onset of the early modern period (and in much of Europe, the Ren ...
, the Western European nobility that used peregrines for hunting, considered the bird associated with
prince A prince is a male ruler (ranked below a king, grand prince, and grand duke) or a male member of a monarch's or former monarch's family. ''Prince'' is also a title of nobility (often highest), often hereditary, in some European states. ...
s in formal hierarchies of birds of prey, just below the gyrfalcon associated with
king King is the title given to a male monarch in a variety of contexts. The female equivalent is queen regnant, queen, which title is also given to the queen consort, consort of a king. *In the context of prehistory, antiquity and contempora ...
s. It was considered "a royal bird, more armed by its courage than its claws". Terminology used by peregrine breeders also used the
Old French Old French (, , ; Modern French: ) was the language spoken in most of the northern half of France from approximately the 8th to the 14th centuries. Rather than a unified language, Old French was a linkage of Romance dialects, mutually intel ...
term , "of noble birth; aristocratic", particularly with the peregrine. The peregrine falcon is the national animal of the
United Arab Emirates The United Arab Emirates (UAE; ar, اَلْإِمَارَات الْعَرَبِيَة الْمُتَحِدَة ), or simply the Emirates ( ar, الِْإمَارَات ), is a country in Western Asia (Middle East, The Middle East). It is ...
. Since 1927, the peregrine falcon has been the official mascot of Bowling Green State University in
Bowling Green, Ohio Bowling Green is a city in and the county seat of Wood County, Ohio, United States, located southwest of Toledo. The population was 30,028 at the 2010 census. It is part of the Toledo Metropolitan Area and a member of the Toledo Metropolitan Ar ...
. The 2007 U.S.
Idaho Idaho ( ) is a U.S. state, state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Monta ...
state quarter features a peregrine falcon. The peregrine falcon has been designated the official city bird of
Chicago (''City in a Garden''); I Will , image_map = , map_caption = Interactive Map of Chicago , coordinates = , coordinates_footnotes = , subdivision_type = List of sovereign states, Count ...
. ''The Peregrine'', by
J. A. Baker John Alec Baker (6 August 1926 – 26 December 1987) was an English author, best known for ''The Peregrine,'' which won the Duff Cooper Prize in 1967. ''The Peregrine'' Robert Macfarlane deemed ''The Peregrine'' to be "a masterpiece of twentie ...
, is widely regarded as one of the best nature books in English written in the twentieth century. Admirers of the book include Robert Macfarlane,
Mark Cocker Mark Cocker (born 1959) is a British author and naturalist. He lives with his wife, Mary Muir, and two daughters in Claxton, Norfolk; the countryside around Claxton is a theme for two of his twelve books. Cocker has written extensively for ...
, who regards the book as "one of the most outstanding books on nature in the twentieth century" and Werner Herzog, who called it "the one book I would ask you to read if you want to make films", and said elsewhere "it has prose of the calibre that we have not seen since
Joseph Conrad Joseph Conrad (born Józef Teodor Konrad Korzeniowski, ; 3 December 1857 – 3 August 1924) was a Polish-British novelist and short story writer. He is regarded as one of the greatest writers in the English language; though he did not spe ...
". In the book, Baker recounts, in diary form, his detailed observations of peregrines (and their interaction with other birds) near his home in
Chelmsford Chelmsford () is a city in the City of Chelmsford district in the county of Essex, England. It is the county town of Essex and one of three cities in the county, along with Southend-on-Sea and Colchester. It is located north-east of Londo ...
, Essex, over a single winter from October to April. An episode of the hour-long TV series '' Starman'' in 1986 titled "Peregrine" was about an injured peregrine falcon and the endangered species program. It was filmed with the assistance of the University of California's peregrine falcon project in Santa Cruz. (Note: the episode was titled "The Falcon" during filming and retitled "Peregrine" before broadcast. An end credit gives thanks to Brian Walton and the Peregrine Fund Facility at UCSC.)


See also

*
List of birds by flight speed This is a list of the fastest flying birds in the world. A bird's velocity is necessarily variable; a hunting bird will reach much greater speeds while diving to catch prey than when flying horizontally. The bird that can achieve the greatest airs ...
*
Perilanner The word perilanner is a falconer's term for a hybrid between a peregrine falcon and a lanner falcon. It is larger and faster than a lanner, but does not fly as far as a peregrine and thus is less likely to fly far away and become lost. As a rule ...
, a hybrid of the peregrine falcon and the lanner falcon (''Falco biarmicus'') * Perlin, a hybrid of the peregrine falcon and the
merlin Merlin ( cy, Myrddin, kw, Marzhin, br, Merzhin) is a mythical figure prominently featured in the legend of King Arthur and best known as a mage, with several other main roles. His usual depiction, based on an amalgamation of historic and leg ...
(''Falco columbarius'')


Explanatory notes


Citations


Sources

* * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * *


Further reading

*


External links

; Conservation organizations
Arctic Raptors – Ongoing research with raptors in the Canadian Arctic

Falcon Research Group

Peregrine Falcon Fund

The Canadian Peregrine Foundation



London Peregrine Partnership (UK)
; Video and other media of peregrines

* ttps://www.youtube.com/watch?v=j3mTPEuFcWk A video of the falcon stooping at a top speed of
Derby Cathedral Peregrine Project, UK. Links to webcams and video sequences

Norwich Cathedral Peregrine Web Cam 2015, UK.
*
The Raptor Resource Project. Links to Peregrine Falcon webcams

Peregrines on Brussels Cathedral


* ttp://www.ntu.ac.uk/ecoweb/biodiversity/falcons/index.html Nottingham Trent University, where peregrines return to breed on the top of the Newton building every year. Includes images and webcam.
University of Massachusetts Amherst Live Falcon Cam at the top of the W.E.B. DuBois library, active each year from when the bonded pair of peregrine falcons brood eggs until the chicks are fledged.

Worcester Peregrine Falcon Project, UK. Includes feeds from 'Peregrines in Worcester' Facebook Fan page, YouTube & Flickr photo groups

Peregrine Falcon Banding
Metropolitan Transportation Authority Bridges and Tunnels; 3 June 2010; 3-minute YouTube video clip
Throgs Neck Bridge Peregrine Banding 2011
Metropolitan Transportation Authority Bridges and Tunnels; 27 May 2011; 10:54 YouTube video clip
Peregrine Falcon Banding 2012
Metropolitan Transportation Authority Bridges and Tunnels; 4 June 2012; 2:40 YouTube video clip
Peregrine Falcon Banding 2016
Metropolitan Transportation Authority Bridges and Tunnels; 2 June 2016; 4:15 YouTube video clip {{DEFAULTSORT:Falcon, peregrine Peregrine falcon Peregrine falcon Cosmopolitan birds Birds of prey of Africa Birds of Asia Birds of Europe Birds of North America Birds of North Africa Birds of South America Diurnal raptors of Australia Peregrine falcon Peregrine falcon Peregrine falcon Birds of the Dominican Republic