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Palomino is a genetic color in
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s, consisting of a gold coat and white mane and tail; the degree of whiteness can vary from bright white to yellow. Genetically, the palomino color is created by a single
allele An allele (, ; ; modern formation from Greek ἄλλος ''állos'', "other") is a variation of the same sequence of nucleotides at the same place on a long DNA molecule, as described in leading textbooks on genetics and evolution. ::"The chrom ...
of a dilution gene called the
cream gene The cream gene is responsible for a number of horse coat colors. Horses that have the cream gene in addition to a base coat color that is chestnut will become palomino if they are heterozygous, having one copy of the cream gene, or cremello, if ...
working on a "red" ( chestnut) base coat. Palomino is created by a genetic mechanism of incomplete dominance, hence it is not considered true-breeding. However, most color breed registries that record palomino horses were founded before equine coat color genetics were understood as well as they are today, therefore the standard definition of a palomino is based on the visible coat color, not heritability nor the underlying presence of the dilution gene. Due to their distinct color, palominos stand out in a show ring, and are much sought after as parade horses. They were particularly popular in movies and television during the 1940s and 1950s. One of the most famous palomino horses was Trigger, known as "the smartest horse in movies", the faithful mount of the Hollywood cowboy star Roy Rogers. Another famous palomino was Mister Ed (real name
Bamboo Harvester Bamboo Harvester (1949–1970) was the American Saddlebred/part-Arabian horse that portrayed Mister Ed on the 1961–1966 comedy series of the same name. Foaled in 1949, the gelding was trained by Will Rogers' protégé, Les Hilton. He was born ...
) who starred on his own TV show in the 1960s. A palomino was also featured in the show '' Xena: Warrior Princess'' (1995–2001). Xena's horse Argo was portrayed by a palomino mare named Tilly. In today's horse breeding the palomino color can be created by crossing a chestnut with a cremello. ''Palomino'' is a Spanish word meaning ''juvenile pigeon'' (the diminutive of ''paloma'', pigeon) and its equine usage refers to the color of such birds.


Description

Palomino horses have a yellow or gold coat, with a white or light cream mane and tail. The shades of the body coat color range from cream to a dark gold. Unless also affected by other, unrelated genes, palominos have dark skin and brown eyes, though some may be born with pinkish skin that darkens with age. Some have slightly lighter brown or amber eyes. A heterozygous cream dilute (CR) such as the palomino must not be confused with a horse carrying champagne dilution. Champagne (CH) dilutes are born with pumpkin-pink skin and blue eyes, which darken within days to amber, green or light brown, and their skin acquires a darker mottled complexion around the eyes, muzzle, and genitalia as the animal matures. A horse with rosy-pink skin and blue eyes in adulthood is most often a cremello or a perlino, a horse carrying two cream dilution genes.
Sooty Sooty is a British children's television media franchise created by Harry Corbett incorporating primarily television and stage shows. The franchise originated with his fictional glove puppet character introduced to television in 1955, with the ...
palomino horses may have darker hairs in the mane, tail and coat. The summer coat of a palomino is usually a slightly darker shade than the winter coat.


Colors confused with palomino

Many non-palominos may also have a gold or tan coat and a light mane and tail. * Chestnut with flaxen mane and tail: Lighter chestnuts with a light cream mane and tail carry a flaxen gene, but not a cream dilution. For example, the Haflinger breed has many light chestnuts with flaxen that may superficially resemble dark palomino, but there is no cream gene in the breed. *
Cremello The cream gene is responsible for a number of horse coat colors. Horses that have the cream gene in addition to a base coat color that is chestnut will become palomino if they are heterozygous, having one copy of the cream gene, or cremello, ...
s carry two copies of the
cream gene The cream gene is responsible for a number of horse coat colors. Horses that have the cream gene in addition to a base coat color that is chestnut will become palomino if they are heterozygous, having one copy of the cream gene, or cremello, if ...
and have a light mane and tail but also a cream-colored hair coat, rosy pink skin and blue eyes. *The champagne gene is the most similar palomino mimic, as it creates a golden-colored coat on some horses, but golden champagnes have light skin with mottling, blue eyes at birth, and amber or hazel eyes in adulthood. *Horses with a very dark brown coat but a flaxen mane and tail are sometimes called "chocolate palomino", and some palomino color registries accept horses of such color. However, this coloring is not genetically palomino. There are two primary ways the color is created. The best-known is a
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it i ...
chestnut with a flaxen mane and tail. The genetics that create light flaxen manes and tails on otherwise chestnut horses are not yet fully understood, but they are not the same as the cream dilution. The other genetic mechanism is derived from the
silver dapple gene The silver or silver dapple (''Z'') gene is a dilution gene that affects the black base coat color and is associated with Multiple Congenital Ocular Abnormalities. It will typically dilute a black mane and tail to a silvery gray or flaxen color ...
, which lightens a black coat to dark brown, and affects the mane and tail even more strongly, diluting to cream or near-white. * Buckskins have a golden body coat but a black mane and tail. Buckskin is also created by the action of a single cream gene, but on a
bay A bay is a recessed, coastal body of water that directly connects to a larger main body of water, such as an ocean, a lake, or another bay. A large bay is usually called a gulf, sea, sound, or bight. A cove is a small, circular bay with a na ...
coat. * Dun horses have a tan body with a darker mane and tail plus primitive markings such as a dorsal stripe down the spine and horizontal striping on the upper back of the forearm. *The pearl gene in a
homozygous Zygosity (the noun, zygote, is from the Greek "yoked," from "yoke") () is the degree to which both copies of a chromosome or gene have the same genetic sequence. In other words, it is the degree of similarity of the alleles in an organism. Mo ...
state creates a somewhat apricot-colored coat with pale skin. When crossed with a single cream gene, the resulting horse, often called a "pseudo-double-dilute", appears visually to be a cremello.


Color breed registries

In the United States, some palomino horses are classified as a color breed. However, unlike the Appaloosa or the Friesian, which are distinct breeds that also happen to have a unique color preference, Palomino color breed registries often accept a wide range of breed or type if the animals are properly golden-colored. The Palomino cannot be a true horse breed, however, because palomino color is an
incomplete dominant In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and t ...
gene and does not breed "true". A palomino crossed with a palomino may result in a palomino about 50% of the time, but could also produce a chestnut (25% probability) or a cremello (25% probability). Thus, palomino is simply a partially expressed color
allele An allele (, ; ; modern formation from Greek ἄλλος ''állos'', "other") is a variation of the same sequence of nucleotides at the same place on a long DNA molecule, as described in leading textbooks on genetics and evolution. ::"The chrom ...
and not a set of characteristics that make up a "breed". Because registration as a palomino with a color breed registry is based primarily on coat color, horses from many breeds or combination of breeds may qualify. Some breeds that have palomino representatives are the American Saddlebred, Tennessee Walking Horse,
Morgan Morgan may refer to: People and fictional characters * Morgan (given name), including a list of people and fictional characters * Morgan le Fay, a powerful witch in Arthurian legend * Morgan (surname), a surname of Welsh origin * Morgan (singer ...
and
Quarter Horse The American Quarter Horse, or Quarter Horse, is an American breed of horse that excels at sprinting short distances. Its name is derived from its ability to outrun other horse breeds in races of a quarter mile or less; some have been clocked at ...
. The color is fairly rare in the Thoroughbred, but does in fact occur and is recognized by The Jockey Club. Some breeds, such as the Haflinger and
Arabian The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plat ...
, may appear to be palomino, but are genetically chestnuts with flaxen manes and tails, as neither breed carries the cream dilution gene. However, in spite of their lack of cream DNA, some palomino color registries have registered such horses if their coat color falls within the acceptable range of shades. While the color standard used by palomino organizations usually describes the ideal body color as that of a "newly minted gold coin" (sometimes mistakenly claimed to be a penny), a wider a body color range is often accepted, ranging from a cream-white color to a deep, dark, chocolate color ("chocolate palomino") that may actually be
silver dapple The silver or silver dapple (''Z'') gene is a dilution gene that affects the black base coat color and is associated with Multiple Congenital Ocular Abnormalities. It will typically dilute a black mane and tail to a silvery gray or flaxen color ...
or liver chestnut with a flaxen mane and tail.


Requirements for registration

In the United States, there are two primary color breed registries for Palomino-colored horses: the Palomino Horse Association (PHA), and the Palomino Horse Breeders of America (PHBA). The Palomino Horse Association (PHA) registers palomino horses of any breed and type "on color and conformation".Registration and Membership Instructions
Palomino Horse Association Registration and Membership, accessed December 6, 2009
The shade of color considered ideal by the PHA is the color of a gold coin, but shades of palomino from light to dark gold are accepted. The mane and tail are required to be white, silver, or ivory, but up to 15% dark or reddish-brown hair is accepted. In the interest of breeding palomino horses, the PHA also registers full double-dilute blue-eyed
cremello The cream gene is responsible for a number of horse coat colors. Horses that have the cream gene in addition to a base coat color that is chestnut will become palomino if they are heterozygous, having one copy of the cream gene, or cremello, ...
s, erroneously called "cremello palominos" by the PHA. Horses that are not recorded by any other registry of unknown pedigree are accepted if their color meets the PHA definition of "palomino". The Palomino Horse Breeders of America (PHBA) has stricter requirements. To be accepted by the PHBA, in addition to color, a horse must have the general structure appropriate to the breeds of light riding type recognized by the PHBA. The adult height of the PHBA horse should be , and the horse must not show
draft horse A draft horse (US), draught horse (UK) or dray horse (from the Old English ''dragan'' meaning "to draw or haul"; compare Dutch ''dragen'' and German ''tragen'' meaning "to carry" and Danish ''drage'' meaning "to draw" or "to fare"), less oft ...
or
pony A pony is a type of small horse ('' Equus ferus caballus''). Depending on the context, a pony may be a horse that is under an approximate or exact height at the withers, or a small horse with a specific conformation and temperament. Compared ...
characteristics. An individual that does not meet the height requirements may still be accepted if it is registered in one of the breed registries recognized by the PHBA. The PHBA usually requires horses or both parents of the horse to be registered by or eligible for registration with certain recognized
breed registries A breed registry, also known as a herdbook, studbook or register, in animal husbandry and the hobby of animal fancy, is an official list of animals within a specific breed whose parents are known. Animals are usually registered by their breeder ...
, including those for the American Quarter Horse, Paint, Appaloosa,
Saddlebred The American Saddlebred is a horse breed from the United States. This breed is referred to as the "Horse America Made". Descended from riding-type horses bred at the time of the American Revolution, the American Saddlebred includes the Narr ...
,
Morgan Morgan may refer to: People and fictional characters * Morgan (given name), including a list of people and fictional characters * Morgan le Fay, a powerful witch in Arthurian legend * Morgan (surname), a surname of Welsh origin * Morgan (singer ...
, Holsteiner,
Arabian The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plat ...
, assorted part-Arabian registries,
Pinto Pinto is a Portuguese, Spanish, Jewish (Sephardic), and Italian surname. It is a high-frequency surname in all Portuguese-speaking countries and is also widely present in Spanish-speaking countries, Italy, India especially in Mangalore, Karnatak ...
(horse division only), Thoroughbred, and assorted gaited horse breeds. Horses with PHBA-registered parents are also eligible even if they are not recorded with any other breed registry. In some situations, mares and geldings may be registered without pedigree on account of their conformation and color only, but stallions must always have pedigrees that are "verified in fact". The ideal PHBA body color is the shade of "a United States gold coin". The mane and tail must be naturally white, and may not have more than 15% black, brown or off-colored hairs. Brown or dark primitive markings are not accepted. PHBA also does not accept horses that are
gray Grey (more common in British English) or gray (more common in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning literally that it is "without color", because it can be compose ...
or show color characteristics of Paints, pintos, Appaloosas or cremellos or perlinos. The skin must be dark, other than pink skin on the face connected to a white marking. The PHBA will not accept a horse for regular registration if it has all three characteristics of a double-dilute cream: light (or pink) skin over the body; white or cream-colored hair over the body; and eyes of a bluish cast. White markings on the face and legs may not exceed certain limits. Leg white may not be higher than the level of the elbow or the stifle, white on the face may not extend past the throatlatch. Spotting and characteristics of the Leopard complex and the various
pinto Pinto is a Portuguese, Spanish, Jewish (Sephardic), and Italian surname. It is a high-frequency surname in all Portuguese-speaking countries and is also widely present in Spanish-speaking countries, Italy, India especially in Mangalore, Karnatak ...
patterns are not accepted, and body spots of less than a 4-inch diameter may be allowed. Horses with non-dark skin on the body, white or creamy coat and pink skin around the eyes are not accepted. Spots of pink skin visible in the muzzle or around the eyes, under the tail and between the hind legs are not accepted. An exception is made for horses registered with the American Saddlebred Horse Association, which may have skin of any color. Accepted eye colors are black, brown, blue and hazel. However, horses with blue or partially blue eyes are accepted only if their registration certificate from a recognized breed association mentions the eye color; they are also accepted on horses of unknown pedigree if they are
gelded A gelding is a castrated male horse or other equine, such as a pony, donkey or a mule. Castration, as well as the elimination of hormonally driven behavior associated with a stallion, allows a male equine to be calmer and better-behaved, making th ...
or spayed.


See also

*
Cream gene The cream gene is responsible for a number of horse coat colors. Horses that have the cream gene in addition to a base coat color that is chestnut will become palomino if they are heterozygous, having one copy of the cream gene, or cremello, if ...
* Dilution gene * Equine coat color * Equine coat color genetics * Palomino rabbit


References


"Horse coat color tests"
from the
UC Davis The University of California, Davis (UC Davis, UCD, or Davis) is a public land-grant research university near Davis, California. Named a Public Ivy, it is the northernmost of the ten campuses of the University of California system. The institu ...
Veterinary Genetics Lab
"Introduction to Coat Color Genetics"
''from'' Veterinary Genetics Laboratory, School of Veterinary Medicine, University of California, Davis. Web Site accessed January 12, 2008


External links


The Palomino Horse Association, founded in 1936Palomino Horse Breeders of America, founded 1941
{{Equine coat colors Horse coat colors Color breeds de:Cream-Gen des Pferdes#Aufhellung beim Fuchs