Pteraspidomorpha is an extinct
class
Class, Classes, or The Class may refer to:
Common uses not otherwise categorized
* Class (biology), a taxonomic rank
* Class (knowledge representation), a collection of individuals or objects
* Class (philosophy), an analytical concept used d ...
of early
jawless fish
Agnatha (; ) or jawless fish is a paraphyletic infraphylum of animals in the subphylum Vertebrata of the phylum Chordata, characterized by the lack of jaws. The group consists of both extant taxon, living (Cyclostomi, cyclostomes such as hagfish ...
. They have long been regarded as closely related or even ancestral to jawed vertebrates, but the few characteristics they share with the latter are now considered as
basal traits for all
vertebrates
Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain.
The vertebrates make up the subphylum Vertebra ...
.
Characteristics

Pteraspidomorphs are characterized by their massive dermal head armour having large, median, ventral and dorsal
plates or shields.
[ Janvier, Philippe (1997]
Pteraspidomorphi
''The Tree of Life Web Project''.
The fossils show extensive shielding of the head. Many had
hypocercal tails in order to generate lift to increase ease of movement through the water for their armoured bodies, which were covered in dermal bone. They also had sucking mouth parts and some species may have lived in fresh water.
Most pteraspidomorphs were marine, but lived very near to the shore, in lagoons and deltas. Some groups are thought to have been fresh water-dwelling. They were certainly bottom-dwellers, as shown by traces of abrasion of the ventral surfaces of their headshields.
Classification
Pteraspidomorphs have been first regarded as related to
bony fish
Osteichthyes ( ; ), also known as osteichthyans or commonly referred to as the bony fish, is a Biodiversity, diverse clade of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondricht ...
es, then to
shark
Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the ...
s, then ancestral to
hagfish
Hagfish, of the Class (biology), class Myxini (also known as Hyperotreti) and Order (biology), order Myxiniformes , are eel-shaped Agnatha, jawless fish (occasionally called slime eels). Hagfish are the only known living Animal, animals that h ...
es, and finally as the closest jawless relatives of the
gnathostomes.
This last theory was based on the fact that they seem to have a paired olfactory organ and a sensory-line pattern which is quite similar to that of the gnathostomes. These characteristics are, however, likely to be general for either the vertebrates or, at any rate, for the ensemble of all
ostracoderms and the gnathostomes. Other ostracoderms, such as the
Galeaspida
Galeaspida (from Latin, 'Helmet shields') is an extinct taxon of jawless marine and freshwater fish. The name is derived from ''galea'', the Latin word for ''helmet'', and refers to their massive bone shield on the head. Galeaspida lived in shallo ...
are now known to have a paired olfactory organ. Current phylogenetic analysis using a large number of characteristics now place pteraspidomorphs as the sister-group of all other ostracoderms and the gnathostomes.
Phylogeny
Based on the work of Mikko Haaramo.
See also
*
Ostracoderm
References
External links
Pteraspidomorphiat Tolweb
at Palaeos
{{Taxonbar, from=Q134673
Early Ordovician first appearances
Late Devonian animals
Late Devonian extinctions
Prehistoric fish classes