Plain Softtail
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The plain softtail (''Thripophaga fusciceps'') is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the Furnariinae subfamily of the ovenbird
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Furnariidae. It is found in
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
,
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
,
Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...
, and
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
.


Taxonomy and systematics

The plain softtail was at times placed by some authors in genus ''
Phacellodomus ''Phacellodomus'' is the genus of thornbirds, birds in the family (biology), family Furnariidae. They are found in woodlands, shrublands, and grasslands, often near water, in South America. Taxonomy The genus ''Phacellodomus'' was described in 1 ...
''. The species has three subspecies, the
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In th ...
''T. f. fusciceps'' ( Sclater, PL, 1889), ''T. f. dimorpha'' ( Bond, J & Meyer de Schauensee, 1941), and ''T. f. obidensis'' (
Todd Todd or Todds may refer to: Places Australia * Todd River, an ephemeral river United States * Todd Valley, California, also known as Todd, an unincorporated community * Todd, Missouri, a ghost town * Todd, North Carolina, an unincorporated c ...
, 1925). Because the subspecies are different sizes and are geographically much separated, some authors speculate that they should be treated as full species.Remsen, Jr., J. V. (2020). Plain Softtail (''Thripophaga fusciceps''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.plasof1.01 retrieved November 11, 2023


Description

The plain softtail is long and weighs . The sexes have almost the same plumage. Adult males of the nominate subspecies have a faint pale
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
and a thin dark line through the eye on a otherwise pale brownish gray face. Their crown is pale gray-brown with faint buff marks. Their upper back is pale gray-brown that becomes rufescent brown on the lower back, rump, and uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
. Their tail is rufous. Their wings are mostly dark rufous with dark brown tips on the flight feathers. Their chin is pale buff-brown and their throat, breast, and belly light brown with some faint paler streaks, and their flanks and undertail coverts are a slightly darker brown. (Some individuals have grayer underparts.) Their iris is highly variable, their
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
bluish horn to pale gray, their
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
pale gray to pale bluish, and their legs and feet olive-green to yellowish olive. Females have paler foreheads and underparts than males. Juveniles are faintly mottled, rather than streaked, on their throat and breast. Subspecies ''T. f. dimorpha'' is significantly smaller than the nominate and rather variable in color. In general it is overall much darker than the nominate, with little or no supercilium, a strongly rufescent back, and underparts that can be more rufescent, browner, or grayer than the nominate's. ''T. f. obidensis'' is even darker brown overall than ''dimorpha'' and has solely brown underparts.


Distribution and habitat

The nominate subspecies of the plain softtail is found in northern Bolivia in the departments of Beni,
La Paz La Paz, officially Nuestra Señora de La Paz (Aymara language, Aymara: Chuqi Yapu ), is the seat of government of the Bolivia, Plurinational State of Bolivia. With 755,732 residents as of 2024, La Paz is the List of Bolivian cities by populati ...
, and
Cochabamba Cochabamba (; ) is a city and municipality in central Bolivia in a valley in the Andes mountain range. It is the capital (political), capital of the Cochabamba Department and the list of cities in Bolivia, fourth largest city in Bolivia, with ...
. Subspecies ''T. f. dimorpha'' is found in separately in eastern Ecuador, northeastern Peru, and southeastern Peru. ''T. f. obidensis'' is found in central Brazil along the lower
Madeira River The Madeira River ( ) is a major waterway in South America. It is estimated to be in length, while the Madeira-Mamoré is estimated near or in length depending on the measuring party and their methods. The Madeira is the biggest tributary of ...
and central Amazon River in the states of Amazonas and
Pará Pará () is a Federative units of Brazil, state of Brazil, located in northern Brazil and traversed by the lower Amazon River. It borders the Brazilian states of Amapá, Maranhão, Tocantins (state), Tocantins, Mato Grosso, Amazonas (Brazilian st ...
. The plain softtail inhabits tropical evergreen and river-edge forest, primarily '' várzea'' and the transition zone between it and dryer landscapes. It does occur in drier deciduous forest. It tends to favor forest edges and areas with dense vine tangles. In elevation it ranges between .


Behavior


Movement

The plain softtail is a year-round resident throughout its range.


Feeding

The plain softtail feeds on
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s. It usually forages in pairs or small groups, often as part of a
mixed-species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...
. It forages at all levels from the forest's understorey to its subcanopy. It gleans its prey from dead leaves, debris, and small branches while acrobatically moving along branches and especially amid vines.


Breeding

The plain softtail's nest is a globe of twigs and soft plant material with two entrance tubes low on the side, placed in a branch fork in the canopy. Nothing else is known about the species' breeding biology.


Vocalization

The song of the plain softtail's nominate subspecies "begins with widely spaced, dry ticking notes, accelerating and rising to extremely high notes, then descends into bursts of sputtering tick notes". That of ''T. f. dimorpha'' is "a sharp, loud, and descending churring". That of ''T. f. obidensis'' is a "very rhythmic chattering with rattles in tune". All three subspecies typically sing in duet.


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the plain softtail as being of Least Concern. It has a large though disjunct range and an unknown population size; the latter is believed to be stable. No immediate threats have been identified. It is considered rare and local.


References

{{Taxonbar, from=Q1259839 plain softtail Birds of the Amazon rainforest Birds of the Ecuadorian Amazon Birds of Peruvian Amazonia Birds of the Bolivian Amazon plain softtail plain softtail Taxonomy articles created by Polbot